| # | Institution | Articles | Citations | R ARank | R CRank |
|---|
| 1 | American Museum of Natural History | 223 | 16,346 | 4 | 3 |
| 2 | Cornell University | 44 | 10,553 | 571 | 115 |
| 3 | University of Michigan | 65 | 9,782 | 683 | 212 |
| 4 | National Scientific and Technical Research Council | 28 | 9,169 | 58 | 1 |
| 5 | Swedish Museum of Natural History | 40 | 7,156 | 16 | 2 |
| 6 | Uppsala University | 23 | 4,491 | 812 | 151 |
| 7 | George Washington University | 39 | 3,901 | 110 | 31 |
| 8 | New York Botanical Garden | 19 | 3,009 | 20 | 8 |
| 9 | Harvard University | 40 | 2,893 | 528 | 538 |
| 10 | National University of Singapore | 14 | 2,701 | 1.7K | 320 |
| 11 | University of California | 31 | 2,636 | 4.1K | 2.2K |
| 12 | Natural History Museum | 66 | 2,624 | 52 | 52 |
| 13 | Brigham Young University | 21 | 2,318 | 180 | 58 |
| 14 | Smithsonian Institution | 29 | 2,205 | 76 | 44 |
| 15 | University of Copenhagen | 35 | 1,896 | 713 | 567 |
| 16 | City College of New York | 1 | 1,774 | 1.5K | 37 |
| 17 | University of Helsinki | 28 | 1,311 | 662 | 590 |
| 18 | University of Texas at Austin | 7 | 1,189 | 4.1K | 1.2K |
| 19 | University of Arizona | 7 | 1,179 | 2.8K | 727 |
| 20 | Umeå University | 6 | 1,126 | 1.3K | 339 |
| 21 | National Center for Scientific Research | 34 | 1,088 | 1.7K | 1.4K |
| 22 | University of Barcelona | 11 | 990 | 1.5K | 544 |
| 23 | University of Melbourne | 17 | 983 | 1.8K | 1.1K |
| 24 | Field Museum of Natural History | 18 | 956 | 39 | 33 |
| 25 | Muséum national d'histoire naturelle | 29 | 894 | 41 | 69 |
| 26 | Chinese Academy of Sciences | 26 | 864 | 2.3K | 1.7K |
| 27 | Commonwealth Scientific and Industrial Research Organisation | 9 | 842 | 468 | 204 |
| 28 | University of British Columbia | 10 | 823 | 3.2K | 1.6K |
| 29 | California Academy of Sciences | 12 | 763 | 26 | 15 |
| 30 | University of Oslo | 7 | 709 | 2.2K | 840 |
| 30 | Finnish Museum of Natural History | 13 | 709 | 8 | 5 |
| 32 | University of Oxford | 10 | 708 | 3.1K | 1.8K |
| 33 | Ohio State University | 20 | 702 | 1.7K | 1.6K |
| 34 | University of São Paulo | 23 | 701 | 1.7K | 1.2K |
| 35 | Stockholm University | 16 | 666 | 583 | 573 |
| 36 | University of Queensland | 8 | 577 | 3.1K | 1.6K |
| 37 | University of Cape Town | 10 | 548 | 906 | 533 |
| 38 | University of Hawaii | 3 | 515 | 1.5K | 385 |
| 39 | University of Gothenburg | 12 | 509 | 1.2K | 1.0K |
| 40 | San Diego State University | 3 | 503 | 1.6K | 338 |
| 40 | Royal Ontario Museum | 7 | 503 | 46 | 28 |
| 42 | Charles University | 7 | 482 | 1.5K | 540 |
| 43 | University of Maryland | 7 | 481 | 3.4K | 1.9K |
| 44 | University of California, Davis | 2 | 473 | 3.0K | 836 |
| 45 | Australian Museum | 6 | 448 | 53 | 24 |
| 46 | University of Bristol | 6 | 440 | 2.6K | 1.4K |
| 47 | (former) Institut National de Recherche pour l'Agriculture, l'Alimentation et l'Environnement | 5 | 434 | 1.1K | 666 |
| 48 | Duke University | 6 | 424 | 2.4K | 1.4K |
| 49 | University of Chicago | 5 | 421 | 3.1K | 1.6K |
| 50 | University of Alberta | 8 | 396 | 3.2K | 2.0K |
| 51 | Industrial University of Santander | 3 | 392 | 250 | 29 |
| 51 | University of California, Riverside | 7 | 392 | 314 | 296 |
| 53 | University of Toronto | 19 | 376 | 2.6K | 3.4K |
| 54 | University of Adelaide | 6 | 371 | 2.1K | 1.2K |
| 55 | University of Wyoming | 3 | 369 | 1.3K | 380 |
| 56 | Auburn University | 4 | 364 | 1.8K | 590 |
| 57 | University of Bonn | 9 | 358 | 1.3K | 1.1K |
| 58 | University of Auckland | 4 | 344 | 3.0K | 1.3K |
| 59 | Stony Brook University | 6 | 340 | 1.4K | 988 |
| 60 | University of Turku | 9 | 330 | 903 | 827 |
| 61 | North Carolina State University | 7 | 328 | 2.0K | 1.4K |
| 62 | National University of Córdoba | 1 | 325 | 1.9K | 176 |
| 62 | National University of Salta | 1 | 325 | 190 | 9 |
| 64 | Iziko Museums of South Africa | 3 | 311 | 29 | 9 |
| 65 | University of Jena | 7 | 310 | 831 | 657 |
| 66 | California State University | 5 | 304 | 2.0K | 788 |
| 67 | Colorado State University | 12 | 301 | 922 | 1.2K |
| 68 | University of Wisconsin–Madison | 7 | 300 | 4.2K | 3.2K |
| 69 | Leiden University | 9 | 297 | 1.0K | 1.1K |
| 70 | Middle Tennessee State University | 12 | 287 | 32 | 53 |
| 71 | Federal University of Rio de Janeiro | 5 | 271 | 2.1K | 830 |
| 71 | California Polytechnic State University | 2 | 271 | 448 | 79 |
| 71 | Shaanxi Normal University | 2 | 271 | 1.1K | 222 |
| 74 | University of California, Berkeley | 3 | 269 | 2.8K | 1.6K |
| 75 | University of Florida | 7 | 268 | 3.9K | 2.9K |
| 76 | Arizona State University | 8 | 252 | 2.0K | 1.9K |
| 77 | University of Cambridge | 6 | 247 | 4.0K | 3.1K |
| 77 | University of Rennes 1 | 4 | 247 | 711 | 402 |
| 79 | University of South Bohemia in České Budějovice | 5 | 245 | 265 | 157 |
| 80 | University of Kentucky | 8 | 244 | 1.9K | 1.8K |
| 81 | British Antarctic Survey | 2 | 242 | 322 | 180 |
| 81 | Natural Environment Research Council | 2 | 242 | 238 | 132 |
| 83 | University of Göttingen | 4 | 239 | 2.1K | 1.3K |
| 84 | Museum of Paleontology Egidio Feruglio | 2 | 238 | 23 | 9 |
| 85 | University of the South Pacific | 2 | 236 | 228 | 29 |
| 86 | China Agricultural University | 6 | 233 | 857 | 634 |
| 87 | University of Central Florida | 2 | 225 | 2.6K | 813 |
| 88 | University of Vienna | 6 | 224 | 2.1K | 1.7K |
| 89 | Natural History Museum Vienna | 4 | 223 | 20 | 8 |
| 90 | State University of New York | 4 | 221 | 3.2K | 1.9K |
| 91 | Swedish University of Agricultural Sciences | 3 | 220 | 1.1K | 737 |
| 92 | Manaaki Whenua – Landcare Research | 3 | 218 | 214 | 143 |
| 92 | Natural History Museum of Bern | 2 | 218 | 24 | 4 |
| 94 | Staatliches Museum für Naturkunde Stuttgart | 2 | 214 | 46 | 13 |
| 95 | Universidade Estadual Paulista | 4 | 213 | 1.7K | 612 |
| 95 | Museo Nacional de Ciencias Naturales | 4 | 213 | 110 | 78 |
| 97 | Florida State University | 3 | 211 | 2.7K | 1.4K |
| 98 | University of Illinois at Urbana–Champaign | 1 | 209 | 6.2K | 2.2K |
| 99 | Natural Resources Canada | 1 | 205 | 716 | 252 |
| 99 | National Academy of Sciences of Ukraine | 1 | 205 | 2.0K | 414 |
| 101 | Lund University | 4 | 199 | 3.6K | 2.4K |
| 102 | University of New South Wales | 7 | 198 | 3.1K | 2.8K |
| 103 | Yale University | 4 | 194 | 3.4K | 2.5K |
| 104 | University of Hamburg | 9 | 193 | 931 | 1.2K |
| 105 | Australian National University | 5 | 191 | 2.4K | 1.9K |
| 106 | University of Alabama | 5 | 190 | 1.9K | 1.4K |
| 106 | Aarhus University | 2 | 190 | 4.6K | 2.1K |
| 108 | Federal University of Rio Grande do Sul | 4 | 185 | 2.4K | 1.1K |
| 108 | Oregon State University | 6 | 185 | 1.6K | 1.6K |
| 110 | Stellenbosch University | 3 | 178 | 1.9K | 940 |
| 111 | University of Akron | 2 | 173 | 1.3K | 504 |
| 112 | University of Reading | 4 | 171 | 1.7K | 1.3K |
| 112 | Palacký University Olomouc | 2 | 171 | 939 | 354 |
| 114 | University of Missouri–St. Louis | 2 | 170 | 69 | 34 |
| 115 | Autonomous University of Madrid | 7 | 169 | 1.1K | 1.1K |
| 115 | Ulm University | 2 | 169 | 1.4K | 643 |
| 115 | New Mexico State University | 2 | 169 | 1.4K | 536 |
| 118 | University of Pretoria | 2 | 164 | 2.5K | 887 |
| 119 | Pontifícia Universidade Católica do Rio Grande do Sul | 2 | 163 | 754 | 152 |
| 120 | Utrecht University | 2 | 160 | 4.7K | 2.7K |
| 121 | Federal University of Minas Gerais | 7 | 156 | 1.3K | 1.1K |
| 122 | Universidade Federal do Paraná | 4 | 153 | 1.5K | 607 |
| 123 | Sun Yat-sen University | 6 | 151 | 2.3K | 1.9K |
| 124 | National Autonomous University of Mexico | 6 | 150 | 2.0K | 1.5K |
| 125 | Australian National Insect Collection | 7 | 148 | 8 | 15 |
| 125 | Université de La Réunion | 1 | 148 | 337 | 79 |
| 127 | University of Arkansas | 1 | 147 | 3.5K | 961 |
| 128 | New York State Museum | 1 | 143 | 90 | 35 |
| 129 | Louisiana State University | 2 | 141 | 3.8K | 1.9K |
| 129 | Natural History Museum of Los Angeles County | 3 | 141 | 65 | 43 |
| 131 | Hokkaido University | 2 | 139 | 4.0K | 1.9K |
| 132 | Rhodes University | 2 | 138 | 786 | 338 |
| 133 | Queen's University Belfast | 1 | 137 | 3.9K | 1.4K |
| 133 | Zoological Society of London | 1 | 137 | 253 | 145 |
| 135 | University of Tübingen | 2 | 136 | 3.4K | 2.0K |
| 136 | Yunnan University | 5 | 135 | 516 | 366 |
| 137 | Medical University of Vienna | 3 | 133 | 1.8K | 1.3K |
| 137 | National University of Colombia | 2 | 133 | 1.5K | 371 |
| 139 | Virginia Institute of Marine Science | 2 | 132 | 218 | 163 |
| 140 | University of Bergen | 4 | 130 | 1.9K | 1.8K |
| 140 | Peking Union Medical College | 2 | 130 | 2.1K | 1.1K |
| 140 | Queensland Museum | 4 | 130 | 38 | 31 |
| 143 | Chongqing Normal University | 2 | 129 | 356 | 119 |
| 144 | Flinders University | 2 | 128 | 2.2K | 1.2K |
| 144 | Midwestern University | 2 | 128 | 285 | 110 |
| 144 | Florida Museum of Natural History | 4 | 128 | 125 | 126 |
| 147 | University of Amsterdam | 5 | 122 | 3.3K | 3.2K |
| 147 | University of Eastern Finland | 2 | 122 | 1.4K | 717 |
| 149 | Tennessee State University | 7 | 121 | 47 | 101 |
| 149 | Kunming Institute of Botany | 7 | 121 | 125 | 191 |
| 151 | Ramapo College | 1 | 120 | 108 | 13 |
| 152 | Université de Montréal | 3 | 119 | 3.7K | 2.8K |
| 153 | Royal Saskatchewan Museum | 2 | 118 | 9 | 5 |
| 154 | Autonomous University of Queretaro | 2 | 117 | 337 | 100 |
| 155 | Michigan State University | 4 | 116 | 4.3K | 3.7K |
| 155 | Carnegie Museum of Natural History | 4 | 116 | 34 | 42 |
| 157 | South Australian Museum | 2 | 113 | 96 | 59 |
| 158 | Kunming Institute of Zoology | 1 | 112 | 154 | 58 |
| 159 | University of the Ryukyus | 2 | 110 | 1.2K | 519 |
| 160 | Georgia Southern University | 2 | 109 | 723 | 284 |
| 160 | Sam Houston State University | 2 | 109 | 450 | 191 |
| 162 | Sichuan University | 3 | 108 | 3.4K | 2.2K |
| 163 | Seattle University | 1 | 107 | 492 | 116 |
| 163 | Ocean University of China | 1 | 107 | 1.8K | 751 |
| 163 | University of Alcalá | 1 | 107 | 2.3K | 768 |
| 163 | Virginia Commonwealth University | 1 | 107 | 4.7K | 2.1K |
| 163 | Catholic University of the North | 1 | 107 | 594 | 153 |
| 168 | Idaho State University | 1 | 106 | 962 | 279 |
| 169 | University of Plymouth | 1 | 105 | 2.3K | 941 |
| 169 | National Tropical Botanical Garden | 2 | 105 | 17 | 14 |
| 171 | University of Buenos Aires | 4 | 103 | 2.0K | 1.5K |
| 172 | University of North Carolina at Charlotte | 8 | 99 | 298 | 782 |
| 173 | University of Connecticut | 1 | 98 | 6.0K | 2.8K |
| 174 | University of Kansas | 3 | 97 | 3.3K | 2.6K |
| 174 | Centre d'Écologie Fonctionnelle et Évolutive | 2 | 97 | 111 | 75 |
| 176 | Polish Academy of Sciences | 3 | 96 | 2.9K | 2.2K |
| 177 | University of Tennessee at Chattanooga | 2 | 95 | 197 | 112 |
| 178 | Freie Universität Berlin | 2 | 94 | 3.1K | 2.1K |
| 179 | Nankai University | 4 | 93 | 1.3K | 1.2K |
| 180 | Plantentuin Meise | 1 | 92 | 94 | 42 |
| 181 | University of Milan | 4 | 90 | 3.2K | 3.1K |
| 181 | Ruhr University Bochum | 2 | 90 | 2.9K | 1.9K |
| 181 | Kyoto University | 1 | 90 | 7.1K | 3.5K |
| 181 | University of Delhi | 1 | 90 | 3.1K | 1.1K |
| 181 | University of Vermont | 6 | 90 | 970 | 1.6K |
| 181 | Federal University of São Paulo | 2 | 90 | 2.2K | 1.2K |
| 187 | Ludwig Maximilian University of Munich | 1 | 89 | 5.7K | 3.1K |
| 188 | University of Pennsylvania | 4 | 88 | 5.0K | 4.9K |
| 189 | University of Washington | 2 | 86 | 8.1K | 5.9K |
| 189 | University of Campinas | 2 | 86 | 3.4K | 1.8K |
| 191 | Autonomous University of Barcelona | 1 | 84 | 5.0K | 2.4K |
| 192 | Bavarian State Collection of Zoology | 2 | 83 | 37 | 34 |
| 192 | University of Oulu | 1 | 83 | 3.4K | 1.7K |
| 194 | University of Guelph | 3 | 81 | 2.2K | 2.1K |
| 195 | Royal Alberta Museum | 4 | 80 | 7 | 18 |
| 196 | University of Ljubljana | 2 | 79 | 2.9K | 1.9K |
| 197 | Tel Aviv University | 2 | 78 | 5.4K | 3.8K |
| 197 | University of La Laguna | 1 | 78 | 1.8K | 715 |
| 199 | University of Alicante | 1 | 77 | 2.2K | 859 |
| 199 | University of Chinese Academy of Sciences | 5 | 77 | 1.6K | 1.6K |
| 201 | Massey University | 1 | 76 | 3.0K | 1.5K |
| 202 | Wayne State University | 1 | 75 | 5.2K | 2.7K |
| 202 | Wageningen University & Research | 4 | 75 | 1.6K | 2.0K |
| 204 | University of Strasbourg | 1 | 74 | 2.3K | 1.2K |
| 205 | University of Bayreuth | 2 | 72 | 1.3K | 1.1K |
| 205 | York University | 2 | 72 | 3.1K | 2.2K |
| 205 | Australian National Herbarium | 2 | 72 | 10 | 10 |
| 208 | University of Rostock | 1 | 71 | 2.4K | 1.3K |
| 208 | University of Padua | 3 | 71 | 4.2K | 3.9K |
| 208 | Chungnam National University | 3 | 71 | 1.3K | 1.1K |
| 208 | Naturalis Biodiversity Center | 5 | 71 | 71 | 107 |
| 208 | Rice University | 2 | 71 | 2.2K | 1.8K |
| 208 | Kōchi University | 1 | 71 | 1.1K | 471 |
| 208 | Tokai University | 1 | 71 | 1.8K | 701 |
| 208 | University of North Dakota | 2 | 71 | 895 | 724 |
| 208 | Mie University | 1 | 71 | 1.5K | 650 |
| 217 | Duquesne University | 1 | 70 | 1.0K | 442 |
| 217 | Montclair State University | 1 | 70 | 975 | 395 |
| 217 | Edinboro University of Pennsylvania | 2 | 70 | 41 | 23 |
| 220 | National University of Tucumán | 2 | 69 | 194 | 162 |
| 220 | Western Australian Museum | 2 | 69 | 81 | 71 |
| 222 | National Museum of Nature and Science | 3 | 68 | 98 | 84 |
| 223 | University of California, San Diego | 5 | 67 | 2.5K | 3.6K |
| 224 | Seoul National University | 2 | 66 | 4.6K | 3.5K |
| 224 | Florida International University | 1 | 66 | 3.6K | 1.8K |
| 226 | Moravian Museum | 1 | 65 | 47 | 15 |
| 226 | Tianjin University of Traditional Chinese Medicine | 1 | 65 | 407 | 208 |
| 226 | Yangtze University | 1 | 65 | 857 | 332 |
| 229 | University of Glasgow | 3 | 64 | 3.9K | 3.7K |
| 229 | Fukuyama University | 1 | 64 | 272 | 136 |
| 231 | Austral University of Chile | 1 | 63 | 1.0K | 460 |
| 231 | Universidad Nacional del Comahue | 1 | 63 | 380 | 178 |
| 233 | University of Louisiana at Lafayette | 1 | 62 | 1.1K | 488 |
| 233 | James Cook University | 2 | 62 | 1.5K | 1.4K |
| 233 | University of Southern Mississippi | 1 | 62 | 1.6K | 782 |
| 236 | Sapienza University of Rome | 2 | 61 | 4.8K | 3.7K |
| 236 | University of Georgia | 1 | 61 | 6.2K | 3.7K |
| 236 | Royal Botanic Gardens | 4 | 61 | 165 | 235 |
| 239 | Czech Academy of Sciences | 3 | 60 | 2.1K | 2.0K |
| 240 | Jagiellonian University | 2 | 58 | 2.5K | 1.7K |
| 240 | University of Sydney | 2 | 58 | 7.3K | 5.9K |
| 242 | University of Sussex | 1 | 57 | 3.6K | 2.3K |
| 242 | Capital Normal University | 3 | 57 | 542 | 523 |
| 244 | University of Colorado | 3 | 56 | 3.7K | 4.0K |
| 244 | National Museum of Marine Biology and Aquarium | 1 | 56 | 70 | 34 |
| 246 | Imperial College London | 2 | 54 | 5.8K | 5.1K |
| 247 | University of Siena | 1 | 53 | 3.0K | 2.0K |
| 247 | Agricultural Research Service | 2 | 53 | 952 | 1.0K |
| 249 | University of Antwerp | 1 | 52 | 3.9K | 2.5K |
| 250 | Federal University of Pelotas | 1 | 51 | 1.8K | 651 |
| 250 | State University of Santa Cruz | 3 | 51 | 229 | 189 |
| 252 | Scripps Institution of Oceanography | 3 | 50 | 234 | 386 |
| 253 | Virginia Tech | 2 | 49 | 4.4K | 3.9K |
| 253 | National Museums of Kenya | 1 | 49 | 184 | 135 |
| 255 | University of North Carolina | 1 | 46 | 6.6K | 4.7K |
| 255 | Universidad de Antioquia | 2 | 46 | 1.0K | 729 |
| 255 | J. Craig Venter Institute | 2 | 46 | 136 | 259 |
| 258 | University of Missouri | 1 | 45 | 6.7K | 4.5K |
| 258 | College of William & Mary | 1 | 45 | 477 | 393 |
| 258 | Federal University of Juiz de Fora | 1 | 45 | 1.6K | 520 |
| 258 | Federal University of Western Pará | 1 | 45 | 229 | 55 |
| 258 | Trinity College | 1 | 45 | 2.7K | 1.9K |
| 263 | University of Algarve | 1 | 44 | 1.1K | 781 |
| 263 | Taiwan Forestry Research Institute | 4 | 44 | 18 | 65 |
| 263 | Pontificia Universidad Católica de Valparaíso | 1 | 44 | 298 | 82 |
| 266 | Complutense University of Madrid | 2 | 43 | 4.3K | 3.4K |
| 266 | University of Texas at Arlington | 2 | 43 | 1.5K | 1.6K |
| 266 | French Agricultural Research Centre for International Development | 1 | 43 | 745 | 563 |
| 269 | King's College London | 2 | 42 | 4.4K | 4.1K |
| 270 | Southeastern Louisiana University | 6 | 41 | 45 | 209 |
| 270 | University of Naples Federico II | 1 | 41 | 4.9K | 3.6K |
| 270 | Tokyo Metropolitan University | 2 | 41 | 1.4K | 1.1K |
| 273 | Universidad Nacional de La Plata | 4 | 40 | 793 | 1.1K |
| 274 | RWTH Aachen University | 1 | 39 | 4.3K | 3.2K |
| 274 | University of Neuchâtel | 1 | 39 | 1.1K | 912 |
| 274 | Syracuse University | 1 | 39 | 3.4K | 2.4K |
| 277 | National Tsing Hua University | 1 | 38 | 2.4K | 1.7K |
| 277 | National Institutes of Health | 1 | 38 | 4.9K | 4.1K |
| 277 | New York Institute of Technology College of Osteopathic Medicine | 1 | 38 | 147 | 93 |
| 277 | United States National Library of Medicine | 1 | 38 | 421 | 391 |
| 277 | National Museum Wales | 1 | 38 | 85 | 67 |
| 282 | Corporación para Investigaciones Biológicas | 1 | 37 | 47 | 52 |
| 283 | Villanova University | 1 | 36 | 1.3K | 887 |
| 284 | City University of New York | 2 | 35 | 3.5K | 3.4K |
| 284 | University of Mississippi | 1 | 35 | 2.1K | 1.5K |
| 284 | Canadian Food Inspection Agency | 1 | 35 | 179 | 165 |
| 287 | State University of Feira de Santana | 1 | 34 | 551 | 251 |
| 287 | Pontifical Xavierian University | 1 | 34 | 1.0K | 522 |
| 287 | Federal University of Paraíba | 1 | 34 | 2.0K | 931 |
| 287 | Okinawa Institute of Science and Technology | 3 | 34 | 155 | 301 |
| 291 | Brown University | 1 | 33 | 4.8K | 3.9K |
| 292 | Aalborg University | 1 | 32 | 2.9K | 2.3K |
| 292 | University of Sheffield | 1 | 32 | 6.6K | 5.2K |
| 292 | King Abdullah University of Science and Technology | 2 | 32 | 967 | 1.2K |
| 292 | Hebei GEO University | 2 | 32 | 108 | 113 |
| 292 | Università degli Studi di Milano Bicocca | 2 | 32 | 1.3K | 1.7K |
| 297 | University of Potsdam | 1 | 31 | 2.0K | 1.7K |
| 297 | University of Toulouse | 1 | 31 | 2.8K | 2.2K |
| 297 | National University of Misiones | 1 | 31 | 88 | 65 |
| 297 | Smithsonian Tropical Research Institute | 1 | 31 | 391 | 401 |
| 297 | University of Hawaii at Manoa | 2 | 31 | 1.5K | 2.1K |
| 302 | University of Southampton | 1 | 30 | 6.1K | 5.0K |
| 302 | Humboldt University of Berlin | 2 | 30 | 3.4K | 3.8K |
| 302 | University of Greifswald | 1 | 30 | 1.6K | 1.4K |
| 302 | Federal University of Lavras | 1 | 30 | 1.1K | 672 |
| 302 | National Oceanography Centre | 1 | 30 | 270 | 285 |
| 307 | University of New Hampshire | 2 | 29 | 1.6K | 2.0K |
| 307 | University of Goroka | 1 | 29 | 8 | 6 |
| 307 | Bernice P. Bishop Museum | 2 | 29 | 65 | 91 |
| 310 | Santa Clara University | 1 | 28 | 875 | 737 |
| 311 | University of Zulia | 1 | 27 | 341 | 238 |
| 311 | National Herbarium of Victoria | 1 | 27 | 24 | 34 |
| 311 | Botanischer Garten und Botanisches Museum Berlin | 1 | 27 | 13 | 18 |
| 314 | Russian Academy of Sciences | 2 | 26 | 4.6K | 4.1K |
| 314 | Beijing Forestry University | 1 | 26 | 1.0K | 857 |
| 314 | Hefei University of Technology | 1 | 26 | 1.8K | 1.4K |
| 317 | Universiti Brunei Darussalam | 1 | 25 | 578 | 493 |
| 317 | Ghent University | 1 | 25 | 6.5K | 5.7K |
| 317 | Universidad San Francisco de Quito | 1 | 25 | 288 | 213 |
| 317 | Ludong University | 1 | 25 | 657 | 483 |
| 321 | University of Koblenz - Landau | 1 | 24 | 372 | 398 |
| 321 | University of the West Indies | 1 | 24 | 1.8K | 1.3K |
| 321 | Hebei University | 2 | 24 | 677 | 749 |
| 321 | University of Wales | 1 | 24 | 2.4K | 2.3K |
| 321 | National University of the Northeast | 1 | 24 | 277 | 180 |
| 321 | Federal University of Viçosa | 1 | 24 | 1.8K | 1.2K |
| 321 | Federal Rural University of Rio de Janeiro | 1 | 24 | 1.0K | 541 |
| 328 | McGill University | 1 | 23 | 8.9K | 7.6K |
| 328 | University of Tartu | 1 | 23 | 2.5K | 2.2K |
| 328 | University of the Western Cape | 1 | 23 | 1.4K | 1.2K |
| 328 | Arkansas State University | 1 | 23 | 418 | 393 |
| 328 | Ohio University | 1 | 23 | 2.6K | 2.2K |
| 328 | University of Richmond | 1 | 23 | 694 | 621 |
| 328 | Hangzhou Normal University | 1 | 23 | 1.1K | 1.0K |
| 328 | University of Tennessee | 1 | 23 | 6.2K | 5.3K |
| 328 | University of Iowa | 1 | 23 | 6.5K | 5.5K |
| 337 | Wellesley College | 1 | 22 | 727 | 772 |
| 337 | University of Oklahoma | 1 | 22 | 4.4K | 3.9K |
| 337 | Colgate University | 2 | 22 | 364 | 575 |
| 337 | University of the Philippines Los Baños | 1 | 22 | 153 | 153 |
| 341 | University of Montpellier | 2 | 21 | 1.6K | 2.2K |
| 341 | Baylor University | 1 | 21 | 2.0K | 1.8K |
| 341 | Södertörn University | 4 | 21 | 54 | 343 |
| 344 | Heidelberg University | 1 | 20 | 5.3K | 5.0K |
| 344 | University of Silesia in Katowice | 2 | 20 | 1.0K | 960 |
| 344 | Prince of Songkla University | 2 | 20 | 1.1K | 1.5K |
| 344 | Nanjing Agricultural University | 2 | 20 | 1.2K | 1.5K |
| 344 | United States Government Publishing Office | 2 | 20 | 447 | 922 |
| 344 | Nanchang University | 1 | 20 | 1.9K | 1.8K |
| 344 | Federal University of Rio Grande | 1 | 20 | 881 | 562 |
| 351 | New Mexico Institute of Mining and Technology | 1 | 19 | 522 | 577 |
| 351 | University of L'Aquila | 1 | 19 | 1.1K | 1.3K |
| 351 | Tiangong University | 1 | 19 | 408 | 310 |
| 351 | Tianjin Normal University | 1 | 19 | 641 | 565 |
| 355 | Fort Hays State University | 2 | 18 | 54 | 154 |
| 355 | Victoria University of Wellington | 2 | 18 | 1.7K | 2.5K |
| 355 | University of Palermo | 2 | 18 | 2.2K | 3.2K |
| 355 | University of Warmia and Mazury in Olsztyn | 1 | 18 | 1.2K | 909 |
| 355 | Centre for Australian National Biodiversity Research | 2 | 18 | 24 | 57 |
| 360 | Masaryk University | 2 | 17 | 1.6K | 2.3K |
| 360 | Westerdijk Institute | 1 | 17 | 121 | 176 |
| 360 | Universidade Federal Rural da Amazônia | 1 | 17 | 230 | 138 |
| 363 | National Marine Fisheries Service | 1 | 16 | 460 | 571 |
| 363 | University of Novi Sad | 1 | 16 | 1.9K | 1.7K |
| 365 | University of Edinburgh | 1 | 15 | 7.8K | 7.2K |
| 365 | Northern Michigan University | 1 | 15 | 227 | 262 |
| 365 | Nanjing Normal University | 1 | 15 | 1.9K | 1.8K |
| 365 | Liverpool John Moores University | 1 | 15 | 2.0K | 2.2K |
| 365 | Royal Botanic Garden Edinburgh | 1 | 15 | 130 | 174 |
| 365 | China Jiliang University | 1 | 15 | 693 | 671 |
| 371 | Chulalongkorn University | 2 | 14 | 2.2K | 3.2K |
| 371 | University of Cádiz | 1 | 14 | 1.6K | 1.6K |
| 371 | Bermuda Institute of Ocean Sciences | 2 | 14 | 43 | 110 |
| 371 | Université de Bamenda | 1 | 14 | 366 | 447 |
| 375 | Rio de Janeiro Botanical Garden | 1 | 13 | 48 | 71 |
| 375 | National Science Foundation | 1 | 13 | 433 | 615 |
| 375 | Kasetsart University | 1 | 13 | 1.5K | 1.6K |
| 375 | Federal University of ABC | 2 | 13 | 561 | 905 |
| 379 | University of Freiburg | 1 | 12 | 3.9K | 4.3K |
| 379 | University of Münster | 2 | 12 | 2.9K | 4.4K |
| 379 | Food and Drug Administration | 2 | 12 | 601 | 1.2K |
| 379 | University of Memphis | 1 | 12 | 2.5K | 2.8K |
| 383 | University of Tokyo | 2 | 11 | 6.9K | 8.1K |
| 383 | National Sun Yat-sen University | 1 | 11 | 1.8K | 2.2K |
| 383 | Northern Arizona University | 1 | 11 | 1.6K | 1.8K |
| 383 | Osaka Prefecture University | 1 | 11 | 1.9K | 2.0K |
| 383 | Shanghai Normal University | 1 | 11 | 1.0K | 1.2K |
| 383 | Kyoto Prefectural University | 1 | 11 | 457 | 572 |
| 383 | University of the Philippines Diliman | 1 | 11 | 372 | 502 |
| 383 | Indonesian Institute of Sciences | 1 | 11 | 377 | 397 |
| 383 | Hebei Normal University | 1 | 11 | 745 | 831 |
| 383 | Universidade Federal da Bahia | 1 | 11 | 2.5K | 2.1K |
| 393 | Monash University | 2 | 10 | 6.2K | 8.3K |
| 393 | University of Graz | 1 | 10 | 2.6K | 3.1K |
| 393 | Katholieke Universiteit Leuven | 1 | 10 | 7.2K | 7.7K |
| 393 | George Mason University | 1 | 10 | 3.6K | 4.0K |
| 393 | King Juan Carlos University | 1 | 10 | 1.7K | 1.9K |
| 393 | City College | 1 | 10 | 285 | 484 |
| 399 | Peking University | 2 | 9 | 4.3K | 5.9K |
| 399 | National Taiwan University | 1 | 9 | 5.4K | 5.8K |
| 399 | University of Cincinnati | 1 | 9 | 5.3K | 5.9K |
| 399 | University of Illinois at Chicago | 1 | 9 | 5.8K | 6.6K |
| 399 | Western Carolina University | 1 | 9 | 541 | 717 |
| 399 | University of New Orleans | 1 | 9 | 1.2K | 1.6K |
| 399 | East China Normal University | 1 | 9 | 2.5K | 2.9K |
| 399 | Universidade Federal do Pará | 1 | 9 | 1.7K | 1.7K |
| 399 | Sorbonne Universités | 1 | 9 | 1.3K | 2.0K |
| 399 | Museu Paraense Emílio Goeldi | 1 | 9 | 209 | 274 |
| 409 | University of Toledo | 1 | 8 | 2.4K | 3.1K |
| 409 | Rowland Institute for Science | 1 | 8 | 70 | 147 |
| 409 | Sabancı University | 1 | 8 | 782 | 1.1K |
| 412 | Purdue University | 1 | 7 | 7.4K | 8.2K |
| 412 | Adam Mickiewicz University in Poznań | 1 | 7 | 1.9K | 2.2K |
| 412 | Southwest University | 2 | 7 | 1.6K | 2.6K |
| 412 | Marche Polytechnic University | 1 | 7 | 2.4K | 3.2K |
| 412 | University of Salento | 1 | 7 | 1.6K | 2.2K |
| 412 | University of the Punjab | 1 | 7 | 1.4K | 1.8K |
| 412 | Federal University of Rio Grande do Norte | 1 | 7 | 2.5K | 2.3K |
| 412 | University of Nebraska State Museum | 1 | 7 | 46 | 68 |
| 412 | Central University of Punjab | 1 | 7 | 265 | 431 |
| 421 | Moscow State University | 1 | 6 | 3.8K | 3.9K |
| 421 | New York University | 1 | 6 | 6.1K | 7.0K |
| 421 | University of Belgrade | 1 | 6 | 4.4K | 5.1K |
| 421 | Norwegian University of Science and Technology | 1 | 6 | 4.5K | 5.7K |
| 421 | University of Veterinary Medicine Vienna | 1 | 6 | 469 | 767 |
| 421 | University of Nebraska at Kearney | 1 | 6 | 228 | 414 |
| 427 | Aix-Marseille University | 1 | 5 | 3.0K | 4.0K |
| 427 | Ilia State University | 1 | 5 | 246 | 417 |
| 427 | Hungarian Natural History Museum | 1 | 5 | 161 | 279 |
| 427 | Swiss Federal Railways | 2 | 5 | 5 | 18 |
| 427 | Universidad Regional Amazónica IKIAM | 1 | 5 | 34 | 87 |
| 432 | University of Wrocław | 1 | 3 | 1.9K | 2.4K |
| 432 | Lincoln University | 1 | 3 | 846 | 1.5K |
| 432 | University of Florence | 1 | 3 | 5.1K | 6.7K |
| 432 | Instituto Butantan | 1 | 3 | 473 | 850 |
| 432 | University of Manitoba | 1 | 3 | 5.6K | 7.4K |
| 432 | University of Otago | 1 | 3 | 4.6K | 6.3K |
| 432 | Nova Southeastern University | 1 | 3 | 1.1K | 2.0K |
| 432 | Universidad Tecnológica Indoamérica | 1 | 3 | 60 | 117 |
| 440 | University of Southern California | 1 | 2 | 7.9K | 10.1K |
| 440 | Carnegie Mellon University | 1 | 2 | 3.8K | 5.3K |
| 440 | University of Turin | 1 | 2 | 5.5K | 7.3K |
| 440 | Weber State University | 1 | 2 | 319 | 736 |
| 440 | Universidade Federal de Alagoas | 1 | 2 | 1.2K | 1.6K |
| 440 | Kharazmi University | 1 | 2 | 960 | 1.6K |
| 440 | University of Sucre | 1 | 2 | 108 | 143 |
| 447 | University of Zurich | 1 | 1 | 5.7K | 8.0K |
| 447 | McMaster University | 2 | 1 | 5.1K | 9.2K |
| 447 | Oswaldo Cruz Foundation | 1 | 1 | 1.7K | 2.6K |
| 447 | Saint Mary's College | 1 | 1 | 94 | 234 |
| 451 | Tennessee Technological University | 1 | 0 | 590 | 1.3K |
| 451 | University of Magdalena | 1 | 0 | 194 | 417 |
| 451 | Academia Sinica | 1 | 0 | 3.5K | 5.2K |
| 451 | Museum of New Zealand Te Papa Tongarewa | 1 | 0 | 74 | 154 |
