| # | Institution | Articles | Citations | R ARank | R CRank |
|---|
| 1 | University of Queensland | 2 | 6,524 | 6.8K | 114 |
| 2 | University of California | 38 | 3,174 | 3.6K | 1.9K |
| 3 | University of Tokyo | 23 | 2,672 | 1.9K | 555 |
| 4 | University of Utah | 5 | 1,777 | 3.0K | 413 |
| 5 | University of Washington | 14 | 1,715 | 2.8K | 1.2K |
| 6 | National Institutes of Health | 14 | 1,714 | 1.6K | 791 |
| 7 | Tel Aviv University | 4 | 1,689 | 3.8K | 416 |
| 8 | Russian Academy of Sciences | 28 | 1,636 | 1.0K | 400 |
| 9 | University of Copenhagen | 14 | 1,492 | 1.7K | 692 |
| 10 | University of Oxford | 13 | 1,491 | 2.5K | 1.1K |
| 11 | University of Colorado | 4 | 1,377 | 3.0K | 517 |
| 12 | University of Iowa | 11 | 1,213 | 1.6K | 601 |
| 13 | Okayama University | 6 | 1,146 | 1.0K | 120 |
| 14 | University of Maryland | 4 | 1,135 | 4.9K | 986 |
| 15 | Tokyo University of Agriculture and Technology | 1 | 1,020 | 1.7K | 75 |
| 16 | University of Konstanz | 8 | 1,015 | 501 | 170 |
| 17 | University of Toronto | 8 | 1,003 | 4.9K | 1.8K |
| 18 | University of Wisconsin–Madison | 10 | 1,000 | 3.3K | 1.6K |
| 19 | University of Regensburg | 5 | 970 | 1.1K | 256 |
| 20 | University of Texas Southwestern Medical Center | 1 | 916 | 3.4K | 537 |
| 21 | Punjab Agricultural University | 12 | 914 | 120 | 19 |
| 22 | University at Buffalo | 6 | 858 | 2.1K | 519 |
| 23 | University of Chicago | 5 | 790 | 3.1K | 1.1K |
| 24 | Australian National University | 5 | 750 | 2.4K | 686 |
| 25 | University of Pennsylvania | 8 | 731 | 3.4K | 1.7K |
| 26 | UCL Medical School | 3 | 715 | 711 | 216 |
| 27 | University of Bonn | 7 | 686 | 1.6K | 670 |
| 28 | Indiana University | 8 | 671 | 1.8K | 850 |
| 29 | Tokyo Medical and Dental University | 2 | 667 | 1.8K | 292 |
| 30 | Aarhus University | 3 | 660 | 3.6K | 862 |
| 31 | Karolinska Institute | 3 | 635 | 3.1K | 1.1K |
| 32 | Stockholm University | 4 | 626 | 2.0K | 604 |
| 33 | University of London | 10 | 620 | 1.8K | 1.0K |
| 34 | Washington University School of Medicine | 8 | 618 | 1.4K | 849 |
| 35 | University of Melbourne | 8 | 606 | 3.4K | 1.6K |
| 36 | University of Edinburgh | 9 | 603 | 2.3K | 1.3K |
| 37 | University of Geneva | 6 | 584 | 1.3K | 604 |
| 38 | Union College | 2 | 577 | 268 | 19 |
| 39 | Howard Hughes Medical Institute | 7 | 575 | 470 | 422 |
| 40 | York University | 4 | 565 | 1.9K | 419 |
| 41 | Ohio State University | 7 | 561 | 3.9K | 1.8K |
| 42 | Syracuse University | 3 | 557 | 1.8K | 352 |
| 43 | University of Basel | 7 | 544 | 1.1K | 638 |
| 44 | Dalhousie University | 2 | 542 | 3.6K | 697 |
| 45 | University of Glasgow | 10 | 538 | 1.6K | 1.0K |
| 45 | National Center for Scientific Research | 9 | 538 | 3.8K | 2.2K |
| 47 | Ohio University | 1 | 527 | 2.6K | 179 |
| 48 | Columbia University | 8 | 517 | 3.2K | 2.0K |
| 48 | McGill University | 8 | 517 | 3.1K | 1.7K |
| 50 | University of Kiel | 3 | 511 | 1.0K | 286 |
| 51 | Austral University of Chile | 2 | 508 | 650 | 45 |
| 52 | Swansea University | 1 | 503 | 3.1K | 295 |
| 53 | University of Michigan | 1 | 498 | 10.6K | 2.7K |
| 54 | Stanford University | 2 | 497 | 6.7K | 2.3K |
| 55 | University of Virginia | 7 | 496 | 2.1K | 1.2K |
| 56 | University of Cambridge | 16 | 493 | 2.0K | 2.1K |
| 57 | Heidelberg University | 5 | 488 | 2.5K | 1.1K |
| 58 | King's College | 7 | 473 | 548 | 324 |
| 59 | University of Florida | 8 | 467 | 3.6K | 2.0K |
| 60 | University of Vienna | 5 | 463 | 2.4K | 1.0K |
| 61 | Northwestern University | 4 | 455 | 3.9K | 1.7K |
| 62 | Osaka Medical College | 1 | 454 | 1.0K | 84 |
| 63 | University of Alberta | 3 | 450 | 5.9K | 1.9K |
| 64 | RMIT University | 1 | 449 | 3.4K | 374 |
| 65 | University of Padua | 8 | 441 | 2.3K | 1.3K |
| 66 | University of Potsdam | 3 | 439 | 1.0K | 252 |
| 67 | Rutgers University | 4 | 431 | 3.8K | 1.5K |
| 68 | Cardiff University | 3 | 415 | 3.0K | 930 |
| 69 | California Institute of Technology | 2 | 414 | 2.3K | 977 |
| 70 | Gunma University | 3 | 412 | 909 | 168 |
| 70 | Waseda University | 3 | 412 | 1.7K | 299 |
| 72 | University of Tübingen | 5 | 407 | 2.0K | 982 |
| 73 | Moscow State University | 6 | 403 | 1.6K | 536 |
| 74 | University of Guelph | 1 | 402 | 3.9K | 764 |
| 74 | New York State Department of Health | 1 | 402 | 853 | 180 |
| 76 | Kobe University | 1 | 400 | 3.1K | 336 |
| 77 | University of Salamanca | 1 | 396 | 2.7K | 261 |
| 78 | Harvard University | 3 | 375 | 4.4K | 2.1K |
| 78 | University of Siena | 4 | 375 | 1.3K | 466 |
| 80 | Lund University | 7 | 372 | 2.5K | 1.7K |
| 81 | University of Rochester | 6 | 368 | 1.7K | 1.2K |
| 81 | University of Mississippi Medical Center | 2 | 368 | 1.2K | 288 |
| 83 | Ghent University | 3 | 364 | 4.1K | 1.6K |
| 84 | Princeton University | 5 | 361 | 2.0K | 1.3K |
| 84 | Hebrew University of Jerusalem | 6 | 361 | 1.9K | 1.1K |
| 86 | John Innes Centre | 7 | 357 | 165 | 173 |
| 87 | University College London | 5 | 354 | 4.1K | 2.3K |
| 88 | Kyoto University | 6 | 351 | 3.5K | 1.8K |
| 89 | Free University | 1 | 348 | 1.5K | 244 |
| 90 | University of Alabama at Birmingham | 7 | 347 | 1.8K | 1.2K |
| 91 | Weill Cornell Medicine | 4 | 346 | 1.8K | 955 |
| 92 | Université libre de Bruxelles | 3 | 337 | 2.2K | 833 |
| 93 | Aberystwyth University | 2 | 329 | 883 | 198 |
| 94 | State University of New York | 8 | 324 | 1.8K | 1.5K |
| 95 | University of Georgia | 6 | 317 | 2.5K | 1.5K |
| 96 | University of Pavia | 1 | 314 | 4.0K | 817 |
| 96 | French National Institute of Health and Medical Research | 3 | 314 | 2.0K | 950 |
| 98 | Washington State University | 4 | 307 | 2.3K | 1.2K |
| 99 | Southern Illinois University Carbondale | 2 | 306 | 2.1K | 420 |
| 100 | Hokkaido University | 3 | 295 | 3.3K | 1.2K |
| 101 | Woman's Medical College of Pennsylvania | 1 | 288 | 123 | 15 |
| 102 | Pierre and Marie Curie University | 7 | 286 | 849 | 788 |
| 103 | University of British Columbia | 6 | 285 | 4.6K | 3.2K |
| 103 | University of Agriculture | 1 | 285 | 1.6K | 232 |
| 105 | University of Minnesota | 4 | 283 | 5.9K | 3.3K |
| 105 | Johann Wolfgang Goethe Universität Frankfurt am Main | 1 | 283 | 8 | 1 |
| 107 | Tokyo Metropolitan University | 2 | 280 | 1.4K | 291 |
| 108 | Hiroshima University | 3 | 275 | 2.5K | 796 |
| 109 | Massachusetts Institute of Technology | 4 | 273 | 3.0K | 2.2K |
| 110 | University of Sheffield | 4 | 271 | 3.4K | 1.8K |
| 111 | University of Marburg | 3 | 270 | 1.7K | 815 |
| 112 | University of Kansas | 3 | 268 | 3.3K | 1.4K |
| 113 | Federal University of Rio de Janeiro | 1 | 265 | 5.2K | 855 |
| 114 | German Cancer Research Center | 2 | 264 | 622 | 335 |
| 114 | Instituto Biológico | 2 | 264 | 218 | 51 |
| 116 | University of Texas at Austin | 4 | 262 | 5.8K | 3.2K |
| 116 | University of Manitoba | 5 | 262 | 2.2K | 1.3K |
| 118 | University of Miami | 2 | 260 | 3.9K | 1.4K |
| 118 | Commonwealth Scientific and Industrial Research Organisation | 3 | 260 | 1.1K | 633 |
| 120 | University of Gothenburg | 5 | 259 | 2.4K | 1.7K |
| 120 | Emory University | 2 | 259 | 3.9K | 1.6K |
| 122 | University of Arizona | 5 | 257 | 3.5K | 2.3K |
| 123 | Amherst College | 2 | 255 | 283 | 76 |
| 123 | Oakland University | 3 | 255 | 630 | 185 |
| 125 | Royal Botanic Garden Edinburgh | 1 | 248 | 130 | 47 |
| 126 | University of Zurich | 2 | 246 | 4.3K | 1.8K |
| 127 | Technical University of Munich | 2 | 243 | 4.4K | 1.8K |
| 127 | Royal Holloway, University of London | 4 | 243 | 608 | 376 |
| 129 | University of Pittsburgh | 4 | 238 | 4.5K | 2.7K |
| 129 | Agricultural Research Service | 2 | 238 | 952 | 528 |
| 131 | University of Sydney | 4 | 237 | 5.2K | 3.1K |
| 131 | University of Wisconsin–Parkside | 1 | 237 | 209 | 14 |
| 133 | Oregon Health & Science University | 3 | 236 | 1.7K | 975 |
| 133 | University of Connecticut Health Center | 4 | 236 | 705 | 514 |
| 135 | Johns Hopkins University | 4 | 235 | 4.0K | 2.6K |
| 136 | Natural History Museum | 1 | 233 | 1.1K | 357 |
| 137 | Brown University | 3 | 232 | 2.8K | 1.5K |
| 137 | University of Münster | 4 | 232 | 1.9K | 1.2K |
| 139 | University of Niš | 1 | 231 | 1.1K | 73 |
| 140 | University of New Mexico | 2 | 230 | 3.3K | 1.2K |
| 141 | Cornell University | 7 | 229 | 2.9K | 2.8K |
| 141 | Case Western Reserve University | 2 | 229 | 4.1K | 1.8K |
| 143 | Leipzig University | 3 | 228 | 2.2K | 1.1K |
| 144 | Humboldt University of Berlin | 1 | 227 | 4.7K | 1.3K |
| 145 | University of Würzburg | 4 | 225 | 1.7K | 1.2K |
| 145 | Radboud University Nijmegen | 3 | 225 | 2.5K | 1.4K |
| 147 | Eunice Kennedy Shriver National Institute of Child Health and Human Development | 2 | 221 | 945 | 562 |
| 148 | University of Innsbruck | 2 | 219 | 2.1K | 827 |
| 149 | Chiba University | 3 | 218 | 2.0K | 801 |
| 150 | University of Helsinki | 1 | 217 | 6.6K | 2.2K |
| 151 | Collège de France | 1 | 216 | 609 | 216 |
| 152 | University of Victoria | 2 | 213 | 2.5K | 947 |
| 153 | École Normale Supérieure | 3 | 212 | 758 | 463 |
| 154 | University of Halle-Wittenberg | 1 | 211 | 1.1K | 271 |
| 155 | National Institute of Genetics | 1 | 210 | 425 | 152 |
| 155 | University of Bath | 2 | 210 | 2.5K | 1.1K |
| 157 | University of Amsterdam | 4 | 206 | 3.7K | 2.5K |
| 157 | University of Hawaii | 3 | 206 | 1.5K | 873 |
| 159 | Osaka University | 3 | 204 | 3.4K | 1.7K |
| 160 | University of Chile | 2 | 202 | 3.0K | 960 |
| 160 | Kumamoto University | 3 | 202 | 1.4K | 590 |
| 162 | Nara Institute of Science and Technology | 2 | 201 | 600 | 227 |
| 162 | University of Murcia | 2 | 201 | 2.0K | 593 |
| 162 | University of California, San Francisco | 4 | 201 | 2.1K | 1.6K |
| 165 | USSR Academy of Medical Sciences | 5 | 200 | 160 | 142 |
| 166 | Tohoku University | 4 | 199 | 3.0K | 1.6K |
| 166 | Colorado State University | 2 | 199 | 3.5K | 1.6K |
| 168 | The Rockefeller University | 4 | 198 | 651 | 656 |
| 169 | National Research Council Canada | 2 | 195 | 1.3K | 688 |
| 170 | Utrecht University | 3 | 194 | 3.8K | 2.4K |
| 170 | National Autonomous University of Mexico | 2 | 194 | 4.1K | 1.2K |
| 170 | University of Bern | 4 | 194 | 2.5K | 1.8K |
| 170 | Leonard M. Miller School of Medicine | 3 | 194 | 1.7K | 1.0K |
| 174 | Hungarian Academy of Sciences | 2 | 193 | 2.2K | 949 |
| 175 | Toho University | 1 | 192 | 1.3K | 218 |
| 176 | University of Wyoming | 2 | 191 | 1.8K | 672 |
| 177 | University of Burgundy | 1 | 189 | 1.5K | 424 |
| 177 | Florida State University | 4 | 189 | 2.2K | 1.5K |
| 179 | Weizmann Institute of Science | 3 | 188 | 1.2K | 830 |
| 180 | Autonomous University of Madrid | 2 | 187 | 2.8K | 1.1K |
| 181 | New England Biolabs | 1 | 186 | 81 | 40 |
| 182 | Michigan State University | 4 | 183 | 4.3K | 2.9K |
| 183 | Oregon National Primate Research Center | 1 | 182 | 340 | 139 |
| 184 | Bielefeld University | 2 | 177 | 1.7K | 682 |
| 185 | Université catholique de Louvain | 1 | 176 | 4.5K | 1.6K |
| 185 | Yeshiva University | 3 | 176 | 583 | 441 |
| 185 | Umeå University | 2 | 176 | 2.9K | 1.4K |
| 188 | Université de Paris | 2 | 175 | 1.1K | 330 |
| 189 | SUNY Downstate Medical Center | 1 | 171 | 1.1K | 298 |
| 190 | National Institute for Basic Biology, Japan | 2 | 168 | 243 | 165 |
| 191 | Oak Ridge National Laboratory | 2 | 167 | 1.8K | 1.1K |
| 192 | Kagoshima University | 1 | 166 | 2.2K | 518 |
| 193 | San Diego State University | 1 | 165 | 3.2K | 983 |
| 193 | Queensland University of Technology | 1 | 165 | 4.2K | 1.4K |
| 195 | Royal London Hospital | 2 | 164 | 613 | 245 |
| 196 | Louis Pasteur University | 2 | 163 | 856 | 527 |
| 197 | Hitotsubashi University | 2 | 162 | 236 | 61 |
| 198 | Charles University | 1 | 161 | 4.6K | 1.4K |
| 198 | Kyoto Prefectural University | 1 | 161 | 457 | 101 |
| 200 | Jawaharlal Nehru University | 1 | 158 | 1.6K | 348 |
| 201 | University of Cologne | 3 | 155 | 2.7K | 1.7K |
| 202 | Yokohama City University | 2 | 153 | 938 | 380 |
| 202 | (former) Institut National de Recherche pour l'Agriculture, l'Alimentation et l'Environnement | 1 | 153 | 2.4K | 1.2K |
| 202 | University of California, Berkeley | 4 | 153 | 2.3K | 2.2K |
| 205 | Yale University | 2 | 151 | 5.0K | 2.8K |
| 205 | Muséum national d'histoire naturelle | 2 | 151 | 659 | 376 |
| 205 | Tenon Hospital | 1 | 151 | 573 | 216 |
| 208 | Lariboisière Hospital | 2 | 150 | 469 | 285 |
| 209 | Odense University | 2 | 149 | 443 | 299 |
| 209 | Roswell Park Comprehensive Cancer Center | 1 | 149 | 845 | 400 |
| 211 | Osaka City University | 3 | 147 | 1.0K | 485 |
| 212 | King's College London | 3 | 146 | 3.5K | 2.3K |
| 212 | Tokyo Institute of Technology | 2 | 146 | 2.7K | 1.3K |
| 212 | National Polytechnic Institute | 1 | 146 | 2.3K | 429 |
| 215 | North Carolina State University | 3 | 145 | 3.5K | 2.4K |
| 216 | Leiden University | 2 | 144 | 3.1K | 1.7K |
| 216 | University of North Carolina at Chapel Hill | 2 | 144 | 4.4K | 2.6K |
| 216 | University of Calcutta | 2 | 144 | 1.3K | 405 |
| 219 | University of Sussex | 1 | 143 | 3.6K | 1.4K |
| 219 | Asia University | 1 | 143 | 651 | 136 |
| 221 | University of California, Santa Cruz | 1 | 141 | 1.1K | 468 |
| 222 | University College Dublin | 2 | 138 | 2.9K | 1.5K |
| 223 | Pennsylvania Hospital | 1 | 137 | 303 | 95 |
| 224 | University of Texas Medical Branch | 2 | 136 | 2.2K | 1.3K |
| 225 | Bordeaux Segalen University | 1 | 135 | 197 | 93 |
| 226 | University of Birmingham | 3 | 134 | 4.5K | 3.0K |
| 226 | RIKEN | 1 | 134 | 1.5K | 591 |
| 226 | Mississippi State University | 2 | 134 | 1.9K | 873 |
| 229 | National Institute of Agrobiological Sciences | 3 | 133 | 235 | 211 |
| 230 | Nagoya University | 5 | 132 | 2.2K | 1.8K |
| 230 | VU University Medical Center | 1 | 132 | 1.8K | 995 |
| 232 | Nihon University | 1 | 131 | 3.0K | 772 |
| 232 | Women's and Children's Hospital | 1 | 131 | 326 | 153 |
| 234 | Pennsylvania State University | 3 | 130 | 6.1K | 4.1K |
| 235 | University of Texas MD Anderson Cancer Center | 1 | 129 | 2.4K | 1.2K |
| 236 | University of Hamburg | 3 | 128 | 2.4K | 1.6K |
| 236 | University Hospital Complex of Granada | 1 | 128 | 276 | 50 |
| 238 | Ludwig Maximilian University of Munich | 3 | 127 | 3.5K | 2.6K |
| 239 | Simon Fraser University | 2 | 126 | 3.1K | 1.7K |
| 240 | University of Adelaide | 3 | 125 | 3.4K | 2.5K |
| 240 | Boston University | 2 | 125 | 4.1K | 2.4K |
| 240 | Western General Hospital | 1 | 125 | 1.0K | 520 |
| 243 | University of Lausanne | 2 | 124 | 2.3K | 1.5K |
| 243 | Sophia University | 1 | 124 | 792 | 186 |
| 243 | Université de Sherbrooke | 2 | 124 | 2.0K | 1.1K |
| 246 | University of Turin | 2 | 123 | 4.1K | 2.3K |
| 246 | Whitehead Institute | 1 | 123 | 270 | 218 |
| 246 | La Trobe University | 4 | 123 | 1.7K | 1.5K |
| 249 | University of Nottingham | 3 | 119 | 4.1K | 3.0K |
| 250 | National Academy of Sciences of Ukraine | 2 | 117 | 1.4K | 638 |
| 250 | University of Idaho | 1 | 117 | 2.1K | 805 |
| 252 | University of Delhi | 5 | 116 | 1.1K | 885 |
| 253 | Goethe University Frankfurt | 1 | 114 | 3.8K | 1.6K |
| 253 | University of Colorado Boulder | 1 | 114 | 3.2K | 1.5K |
| 255 | Chonnam National University | 2 | 113 | 1.6K | 866 |
| 256 | Eppley Institute for Research in Cancer and Allied Diseases | 1 | 112 | 84 | 47 |
| 256 | University of Nebraska Medical Center | 1 | 112 | 2.7K | 1.2K |
| 258 | Clark University | 1 | 111 | 960 | 405 |
| 259 | University of Hong Kong | 2 | 110 | 5.1K | 3.1K |
| 260 | University of Milan | 2 | 109 | 4.6K | 2.9K |
| 260 | Katholieke Universiteit Leuven | 2 | 109 | 5.4K | 3.5K |
| 262 | University of North Carolina | 2 | 107 | 5.0K | 3.3K |
| 263 | University of Giessen | 3 | 106 | 1.8K | 1.5K |
| 263 | MRC Human Genetics Unit | 1 | 106 | 60 | 43 |
| 263 | University of Hyogo | 2 | 106 | 808 | 353 |
| 266 | University of Missouri | 1 | 104 | 6.7K | 3.1K |
| 266 | Japan Science and Technology Agency | 2 | 104 | 660 | 530 |
| 268 | University of Oregon | 1 | 103 | 3.2K | 1.5K |
| 268 | Loma Linda University | 1 | 103 | 1.5K | 670 |
| 268 | University of Kentucky | 1 | 103 | 6.3K | 2.9K |
| 271 | Carnegie Institution for Science | 1 | 102 | 589 | 393 |
| 272 | Johns Hopkins School of Medicine | 2 | 100 | 2.1K | 1.6K |
| 272 | Duke University Hospital | 2 | 100 | 2.9K | 2.2K |
| 274 | Teikyo University | 1 | 99 | 974 | 330 |
| 274 | Institut Jacques Monod | 2 | 99 | 106 | 111 |
| 276 | University of Göttingen | 1 | 97 | 4.3K | 2.3K |
| 276 | Nara Women's University | 1 | 97 | 521 | 168 |
| 276 | Beckman Laser Institute | 1 | 97 | 149 | 90 |
| 279 | Marquette University | 1 | 93 | 1.9K | 647 |
| 279 | Deakin University | 2 | 93 | 2.9K | 1.9K |
| 281 | University of New South Wales | 1 | 91 | 8.2K | 4.2K |
| 281 | Hvidovre Hospital | 1 | 91 | 566 | 359 |
| 281 | NYU Langone Medical Center | 3 | 91 | 942 | 944 |
| 284 | McMaster University | 1 | 90 | 6.7K | 3.5K |
| 284 | Cincinnati Children's Hospital Medical Center | 1 | 90 | 1.7K | 1.0K |
| 286 | University of Camerino | 1 | 89 | 1.2K | 512 |
| 286 | University of São Paulo | 1 | 89 | 9.4K | 4.2K |
| 286 | Marche Polytechnic University | 1 | 89 | 2.4K | 1.1K |
| 286 | Hadassah Medical Center | 1 | 89 | 966 | 497 |
| 290 | University of Manchester | 2 | 88 | 6.6K | 4.5K |
| 291 | Peter MacCallum Cancer Centre | 1 | 87 | 394 | 223 |
| 291 | Zoological Society of London | 1 | 87 | 253 | 192 |
| 293 | University of Antwerp | 1 | 86 | 3.9K | 1.9K |
| 293 | University of Rouen | 1 | 86 | 376 | 193 |
| 295 | University of Puget Sound | 1 | 85 | 287 | 87 |
| 295 | University of Houston | 2 | 85 | 3.2K | 2.1K |
| 297 | Brooklyn College | 2 | 84 | 436 | 270 |
| 297 | City University of New York | 2 | 84 | 3.5K | 2.2K |
| 297 | Iwate University | 1 | 84 | 1.0K | 364 |
| 297 | Bispebjerg Hospital | 1 | 84 | 703 | 454 |
| 301 | University of Fribourg | 1 | 83 | 1.6K | 782 |
| 301 | University of Modena and Reggio Emilia | 1 | 83 | 3.1K | 1.6K |
| 301 | Novosibirsk State University | 1 | 83 | 1.1K | 374 |
| 304 | University of Hohenheim | 2 | 82 | 1.0K | 830 |
| 304 | Yamaguchi University | 3 | 82 | 951 | 665 |
| 304 | University of Bari | 2 | 82 | 2.0K | 1.5K |
| 304 | National Institute of Environmental Health Sciences | 1 | 82 | 1.1K | 756 |
| 308 | University of Tsukuba | 3 | 81 | 3.0K | 2.2K |
| 308 | University of Illinois at Chicago | 2 | 81 | 4.3K | 3.1K |
| 308 | Mayo Clinic College of Medicine and Science | 1 | 81 | 1.5K | 1.0K |
| 311 | St George's, University of London | 1 | 80 | 916 | 565 |
| 312 | ETH Zürich | 1 | 78 | 4.8K | 3.0K |
| 313 | Kansas State University | 1 | 77 | 3.5K | 1.8K |
| 313 | University of Rennes 1 | 2 | 77 | 1.3K | 1.0K |
| 313 | Ben-Gurion University of the Negev | 1 | 77 | 4.7K | 2.3K |
| 313 | University of South Alabama | 1 | 77 | 1.6K | 741 |
| 317 | Pitié-Salpêtrière Hospital | 1 | 76 | 1.1K | 700 |
| 318 | Center for Infectious Disease Research | 1 | 75 | 110 | 111 |
| 318 | Cancer Research UK | 2 | 75 | 386 | 420 |
| 320 | University of Liège | 2 | 74 | 2.8K | 2.3K |
| 320 | State University of New York Upstate Medical University | 2 | 74 | 1.1K | 850 |
| 322 | Monash University | 2 | 73 | 6.2K | 4.5K |
| 322 | University of California, San Diego | 2 | 73 | 4.2K | 3.5K |
| 324 | University of Bologna | 1 | 72 | 6.9K | 3.8K |
| 324 | Saitama University | 1 | 72 | 1.2K | 485 |
| 324 | National Eye Institute | 1 | 72 | 413 | 333 |
| 324 | Sambalpur University | 1 | 72 | 270 | 68 |
| 328 | Sapienza University of Rome | 1 | 70 | 6.2K | 3.5K |
| 328 | Technion – Israel Institute of Technology | 1 | 70 | 3.8K | 2.2K |
| 328 | Oklahoma State University–Stillwater | 1 | 70 | 218 | 103 |
| 328 | University of L'Aquila | 1 | 70 | 1.1K | 597 |
| 332 | Université du Québec | 1 | 69 | 1.1K | 677 |
| 332 | University of Auckland | 2 | 69 | 4.6K | 3.6K |
| 332 | Iowa State University | 3 | 69 | 3.3K | 3.1K |
| 332 | Hospital Ramos Mejía | 1 | 69 | 37 | 21 |
| 336 | Purdue University | 3 | 68 | 4.7K | 4.3K |
| 336 | University of Leeds | 2 | 68 | 5.2K | 3.9K |
| 336 | University of Miyazaki | 1 | 68 | 1.7K | 758 |
| 336 | National Institute of Dental and Craniofacial Research | 1 | 68 | 625 | 544 |
| 336 | European Neuroscience Institute Göttingen | 1 | 68 | 11 | 16 |
| 341 | Loyola University | 1 | 67 | 993 | 466 |
| 341 | National Hellenic Research Foundation | 1 | 67 | 386 | 206 |
| 341 | Christchurch Hospital | 1 | 67 | 558 | 345 |
| 341 | University of Nebraska–Lincoln | 1 | 67 | 3.3K | 1.9K |
| 345 | Universidade do Porto | 1 | 66 | 2.4K | 1.3K |
| 346 | Leiden University Medical Center | 1 | 65 | 2.7K | 1.9K |
| 347 | Burnet Institute | 1 | 64 | 250 | 199 |
| 347 | LIRMM | 1 | 64 | 194 | 75 |
| 349 | Consiglio Nazionale delle Ricerche | 1 | 63 | 2.0K | 1.3K |
| 349 | Rosenstiel School of Marine and Atmospheric Science | 1 | 63 | 137 | 125 |
| 351 | Medical College of Wisconsin | 3 | 62 | 1.7K | 1.7K |
| 352 | Louisiana State University | 1 | 59 | 5.3K | 3.1K |
| 352 | Netherlands Institute for Neuroscience | 1 | 59 | 268 | 261 |
| 352 | Texas Tech University Health Sciences Center | 1 | 59 | 1.5K | 870 |
| 352 | Huntsman Cancer Institute | 1 | 59 | 495 | 362 |
| 356 | University of Southern California | 1 | 58 | 7.9K | 5.0K |
| 356 | Hospital Universitario de La Princesa | 1 | 58 | 468 | 288 |
| 358 | Thomas Jefferson University | 1 | 56 | 3.1K | 2.0K |
| 359 | Karlsruhe Institute of Technology | 1 | 55 | 3.5K | 2.3K |
| 359 | University of Massachusetts | 1 | 55 | 5.6K | 3.6K |
| 361 | University of Pisa | 1 | 54 | 5.2K | 3.3K |
| 361 | University of Calicut | 1 | 54 | 341 | 135 |
| 363 | University of Belgrade | 4 | 53 | 2.0K | 2.3K |
| 363 | University of Calgary | 2 | 53 | 5.5K | 4.6K |
| 363 | University of Tennessee | 2 | 53 | 4.6K | 3.9K |
| 366 | University of Groningen | 2 | 52 | 4.4K | 4.0K |
| 366 | Serbian Academy of Sciences and Arts | 4 | 52 | 42 | 59 |
| 366 | Baylor College of Medicine | 1 | 52 | 4.2K | 2.9K |
| 366 | Bucknell University | 1 | 52 | 805 | 414 |
| 370 | Ruhr University Bochum | 3 | 51 | 2.3K | 2.6K |
| 370 | Argonne National Laboratory | 1 | 51 | 1.7K | 1.2K |
| 370 | UCSF Medical Center | 1 | 51 | 604 | 430 |
| 370 | United States National Library of Medicine | 1 | 51 | 421 | 348 |
| 374 | Institut Pasteur de Lille | 1 | 50 | 459 | 431 |
| 375 | Seoul National University | 1 | 49 | 5.9K | 3.9K |
| 376 | Battelle Memorial Institute | 1 | 48 | 431 | 315 |
| 376 | Biozentrum University of Basel | 1 | 48 | 107 | 134 |
| 378 | National Institute of Advanced Industrial Science and Technology | 1 | 47 | 1.7K | 1.1K |
| 378 | Griffith University | 1 | 47 | 4.7K | 3.1K |
| 378 | Bangalore University | 1 | 47 | 593 | 283 |
| 381 | Brookhaven National Laboratory | 1 | 46 | 1.3K | 1.1K |
| 381 | Umm Al-Qura University | 1 | 46 | 977 | 457 |
| 383 | University of Düsseldorf | 1 | 45 | 1.9K | 1.5K |
| 383 | University of Poitiers | 1 | 45 | 1.5K | 1.0K |
| 385 | University of the Basque Country | 1 | 44 | 2.9K | 1.7K |
| 386 | Trinity University | 1 | 43 | 525 | 304 |
| 386 | Norwegian University of Life Sciences | 2 | 43 | 834 | 999 |
| 386 | National Institute for Physiological Sciences | 1 | 43 | 326 | 298 |
| 386 | Universidad Autónoma del Estado de Morelos | 1 | 43 | 692 | 329 |
| 390 | Brandeis University | 1 | 42 | 1.6K | 1.3K |
| 390 | Shimane University | 1 | 42 | 1.5K | 820 |
| 392 | Beth Israel Deaconess Medical Center | 1 | 41 | 2.8K | 2.2K |
| 392 | Northeast Ohio Medical University | 1 | 41 | 598 | 463 |
| 394 | Wake Forest University | 2 | 40 | 1.4K | 1.7K |
| 394 | Macquarie University | 1 | 40 | 4.3K | 3.0K |
| 394 | Queen's University Belfast | 1 | 40 | 3.9K | 2.8K |
| 394 | Statens Serum Institut | 1 | 40 | 573 | 559 |
| 398 | University of Massachusetts Amherst | 1 | 38 | 2.6K | 1.9K |
| 398 | Clemson University | 1 | 38 | 3.7K | 2.6K |
| 400 | United States Public Health Service | 1 | 37 | 137 | 122 |
| 400 | University of South Carolina | 1 | 37 | 5.4K | 4.0K |
| 400 | Paris Diderot University | 2 | 37 | 1.6K | 1.8K |
| 400 | University of California, Davis | 2 | 37 | 3.0K | 3.5K |
| 404 | Anschutz Medical Campus | 2 | 36 | 1.4K | 1.7K |
| 404 | University of Ottawa Heart Institute | 1 | 36 | 429 | 395 |
| 406 | City College of New York | 1 | 35 | 1.5K | 1.1K |
| 406 | Taisei Corporation | 1 | 35 | 81 | 43 |
| 408 | University of Arkansas for Medical Sciences | 1 | 34 | 2.5K | 2.0K |
| 408 | National Institute of Allergy and Infectious Diseases | 1 | 34 | 1.1K | 1.1K |
| 410 | University College Cork | 1 | 33 | 2.7K | 2.2K |
| 410 | De Montfort University | 1 | 33 | 1.6K | 1.1K |
| 412 | Technische Universität Darmstadt | 1 | 32 | 2.4K | 2.0K |
| 412 | University of New Brunswick | 1 | 32 | 2.1K | 1.5K |
| 412 | University of Westminster | 1 | 32 | 1.1K | 867 |
| 412 | Imperial College School of Medicine | 1 | 32 | 749 | 798 |
| 412 | University of Cincinnati Academic Health Center | 1 | 32 | 1.9K | 1.7K |
| 417 | Virginia Tech | 1 | 31 | 5.9K | 4.6K |
| 417 | Shizuoka University | 1 | 31 | 1.6K | 1.1K |
| 417 | Occidental College | 1 | 31 | 344 | 290 |
| 417 | State University of New York at Albany | 1 | 31 | 958 | 864 |
| 417 | Chinese University of Hong Kong | 1 | 31 | 5.9K | 4.7K |
| 417 | Michigan Medicine | 2 | 31 | 2.3K | 2.6K |
| 423 | University of Oklahoma | 1 | 30 | 4.4K | 3.4K |
| 423 | Graz University of Technology | 1 | 30 | 1.7K | 1.5K |
| 423 | Tulane University | 2 | 30 | 2.1K | 2.6K |
| 423 | Herlev Hospital | 1 | 30 | 706 | 683 |
| 423 | University of Texas Health Science Center at San Antonio | 1 | 30 | 2.2K | 2.0K |
| 423 | Strong Memorial Hospital | 1 | 30 | 196 | 177 |
| 423 | National Institute of Occupational Health | 1 | 30 | 362 | 372 |
| 430 | University of Alabama | 2 | 29 | 3.7K | 4.0K |
| 431 | University of Buenos Aires | 1 | 28 | 4.5K | 2.8K |
| 431 | Wistar Institute | 1 | 28 | 426 | 502 |
| 433 | New York University | 1 | 27 | 6.1K | 4.8K |
| 433 | University of Extremadura | 1 | 27 | 2.1K | 1.6K |
| 435 | Bowdoin College | 1 | 26 | 470 | 406 |
| 435 | Albert Einstein College of Medicine | 1 | 26 | 2.8K | 2.6K |
| 435 | Loyola University Medical Center | 1 | 26 | 1.1K | 953 |
| 438 | Arizona State University | 1 | 25 | 6.6K | 5.5K |
| 438 | University of Messina | 1 | 25 | 2.9K | 2.5K |
| 438 | University of Catania | 1 | 25 | 3.6K | 3.2K |
| 438 | Allahabad University | 1 | 25 | 109 | 66 |
| 438 | Cancer Research Center | 1 | 25 | 265 | 336 |
| 443 | Hôpital Saint-Louis | 1 | 24 | 564 | 572 |
| 444 | Rice University | 1 | 23 | 3.1K | 3.0K |
| 444 | Université du Québec à Montréal | 1 | 23 | 2.3K | 2.2K |
| 444 | Kansai Medical University | 2 | 23 | 726 | 892 |
| 447 | University of Louisiana at Lafayette | 1 | 22 | 1.1K | 1.0K |
| 447 | University of Louisville | 1 | 22 | 4.1K | 3.6K |
| 449 | Polish Academy of Sciences | 1 | 21 | 4.5K | 3.9K |
| 449 | Clarkson University | 1 | 21 | 1.2K | 1.2K |
| 449 | National University of Ireland | 1 | 21 | 2.4K | 2.5K |
| 449 | New York Medical College | 1 | 21 | 1.7K | 1.6K |
| 449 | Skidmore College | 1 | 21 | 369 | 413 |
| 449 | Simmons University | 1 | 21 | 332 | 322 |
| 455 | Duke University | 1 | 20 | 6.1K | 5.7K |
| 455 | University of Yamanashi | 1 | 20 | 1.6K | 1.4K |
| 455 | University of Exeter | 1 | 20 | 4.4K | 4.1K |
| 455 | National Institute on Aging | 1 | 20 | 961 | 1.2K |
| 459 | Concordia University | 1 | 19 | 3.5K | 3.1K |
| 459 | University of Duisburg-Essen | 1 | 19 | 3.4K | 3.3K |
| 459 | Ulm University | 1 | 19 | 2.0K | 2.1K |
| 459 | Miyagi University of Education | 1 | 19 | 133 | 129 |
| 459 | Medizinische Hochschule Hannover | 1 | 19 | 860 | 957 |
| 464 | University of Coimbra | 1 | 18 | 3.6K | 3.3K |
| 464 | University of Bristol | 1 | 18 | 6.4K | 6.0K |
| 464 | University of Warwick | 1 | 18 | 5.2K | 4.9K |
| 464 | University of Reading | 1 | 18 | 3.7K | 3.6K |
| 464 | Bose Institute | 1 | 18 | 505 | 517 |
| 464 | University of Kalyani | 1 | 18 | 745 | 647 |
| 464 | Southern Illinois University School of Medicine | 1 | 18 | 765 | 916 |
| 471 | Semmelweis University | 1 | 17 | 2.0K | 2.0K |
| 471 | Eli Lilly and Company | 1 | 17 | 1.1K | 1.3K |
| 471 | University of Waterloo | 1 | 17 | 5.6K | 5.3K |
| 471 | Heart Research Institute | 1 | 17 | 72 | 131 |
| 475 | University of Oviedo | 1 | 16 | 2.8K | 2.8K |
| 476 | Shinshu University | 1 | 15 | 2.0K | 1.9K |
| 477 | Binghamton University | 1 | 14 | 1.7K | 1.9K |
| 477 | Oita University | 1 | 14 | 1.4K | 1.3K |
| 477 | University of Paris-Sud | 1 | 14 | 2.1K | 2.5K |
| 477 | John Curtin School of Medical Research | 1 | 14 | 391 | 568 |
| 477 | The Open University of Japan | 1 | 14 | 136 | 130 |
| 482 | Panjab University | 1 | 13 | 1.7K | 1.7K |
| 482 | University of Caen Normandy | 1 | 13 | 897 | 1.3K |
| 482 | Mayo Clinic | 1 | 13 | 4.5K | 4.4K |
| 482 | Edinburgh Napier University | 1 | 13 | 991 | 1.2K |
| 486 | University of Southampton | 1 | 12 | 6.1K | 6.4K |
| 486 | Tottori University | 1 | 12 | 1.8K | 1.9K |
| 486 | University of Parma | 1 | 12 | 2.9K | 3.4K |
| 486 | Southern Oregon University | 1 | 12 | 122 | 188 |
| 490 | University of Pécs | 1 | 11 | 1.5K | 1.8K |
| 490 | Tokyo Dental College | 1 | 11 | 423 | 501 |
| 490 | National Institute of Diabetes and Digestive and Kidney Diseases | 1 | 11 | 501 | 785 |
| 493 | Massachusetts General Hospital | 1 | 10 | 4.5K | 4.9K |
| 493 | Carnegie Mellon University | 1 | 10 | 3.8K | 4.3K |
| 493 | University of Otago | 1 | 10 | 4.6K | 5.2K |
| 493 | University of Algarve | 1 | 10 | 1.1K | 1.5K |
| 493 | Medical University of South Carolina | 1 | 10 | 3.3K | 3.6K |
| 493 | Royal Hallamshire Hospital | 1 | 10 | 848 | 1.1K |
| 493 | Wellcome Genome Campus | 1 | 10 | 391 | 645 |
| 493 | University of Newcastle | 1 | 10 | 5.0K | 5.7K |
| 501 | University of Palermo | 1 | 9 | 3.2K | 3.9K |
| 501 | University of Toledo Medical Center | 1 | 9 | 729 | 986 |
| 501 | Sri Venkateswara College | 1 | 9 | 84 | 142 |
| 501 | Visva Bharati University | 1 | 9 | 101 | 161 |
| 505 | Qatar University | 1 | 8 | 1.9K | 2.5K |
| 506 | Worcester Polytechnic Institute | 2 | 7 | 800 | 1.8K |
| 506 | Brock University | 1 | 7 | 1.7K | 2.4K |
| 506 | University of Tokushima | 1 | 7 | 1.7K | 2.0K |
| 509 | Ochanomizu University | 1 | 6 | 676 | 946 |
| 510 | Addenbrooke's Hospital | 1 | 5 | 1.7K | 2.3K |
| 510 | Adam Mickiewicz University in Poznań | 1 | 5 | 1.9K | 2.4K |
| 510 | Scripps Research | 1 | 5 | 1.5K | 2.1K |
| 510 | California State University | 1 | 5 | 5.5K | 6.4K |
| 510 | University of Calabria | 1 | 5 | 2.0K | 2.8K |
| 510 | University of Maryland Medical Center | 1 | 5 | 635 | 972 |
| 510 | Albert Szent-Györgyi University of Medicine | 1 | 5 | 197 | 331 |
| 517 | Illinois Institute of Technology | 1 | 4 | 1.6K | 2.5K |
| 517 | Wye College | 2 | 4 | 100 | 217 |
| 519 | University of Bremen | 1 | 3 | 2.6K | 4.0K |
| 519 | Cajal Institute | 1 | 3 | 82 | 177 |
| 519 | Montefiore Hospital | 1 | 3 | 177 | 329 |
| 522 | Nippon Steel | 1 | 1 | 300 | 495 |
| 522 | University of Richmond | 1 | 1 | 694 | 1.4K |
| 522 | University of Texas System | 1 | 1 | 2.4K | 3.7K |
| 522 | University of Montana | 1 | 1 | 1.5K | 2.7K |
| 526 | Freie Universität Berlin | 1 | 0 | 4.4K | 6.8K |
| 526 | Newcastle University | 1 | 0 | 5.4K | 8.3K |
| 526 | Royal Victoria Infirmary | 1 | 0 | 661 | 1.2K |
