| # | Institution | Articles | Citations | R ARank | R CRank |
|---|
| 1 | Howard Hughes Medical Institute | 122 | 14,809 | 62 | 65 |
| 2 | University of California | 161 | 12,877 | 966 | 612 |
| 3 | National Institutes of Health | 72 | 6,010 | 415 | 332 |
| 4 | Cancer Research UK | 34 | 4,907 | 41 | 27 |
| 5 | University of Pennsylvania | 44 | 4,493 | 806 | 416 |
| 6 | University of Cambridge | 53 | 4,018 | 662 | 441 |
| 7 | National Center for Scientific Research | 49 | 3,556 | 1.2K | 552 |
| 8 | University of California, San Diego | 25 | 3,381 | 658 | 293 |
| 9 | University of Edinburgh | 31 | 3,284 | 742 | 290 |
| 10 | Baylor College of Medicine | 31 | 2,928 | 569 | 263 |
| 11 | Harvard University | 30 | 2,510 | 734 | 599 |
| 12 | Johns Hopkins School of Medicine | 12 | 2,457 | 615 | 172 |
| 13 | University of Chicago | 34 | 2,337 | 673 | 509 |
| 14 | Memorial Sloan Kettering Cancer Center | 33 | 2,193 | 330 | 248 |
| 15 | University of Texas Southwestern Medical Center | 17 | 2,143 | 691 | 281 |
| 16 | University of Dundee | 20 | 2,135 | 235 | 116 |
| 17 | The Rockefeller University | 20 | 2,086 | 215 | 169 |
| 18 | Ludwig Maximilian University of Munich | 14 | 1,975 | 1.2K | 356 |
| 19 | California Institute of Technology | 24 | 1,956 | 494 | 360 |
| 20 | Columbia University | 23 | 1,878 | 1.4K | 759 |
| 21 | University of Washington | 25 | 1,854 | 1.7K | 1.1K |
| 22 | Wistar Institute | 5 | 1,845 | 130 | 36 |
| 23 | University of Geneva | 19 | 1,817 | 386 | 198 |
| 24 | Pasteur Institute | 15 | 1,806 | 336 | 168 |
| 25 | University of Oxford | 42 | 1,776 | 866 | 936 |
| 26 | University College London | 22 | 1,689 | 1.3K | 755 |
| 27 | Scripps Research | 11 | 1,621 | 384 | 222 |
| 28 | Beth Israel Deaconess Medical Center | 9 | 1,617 | 858 | 291 |
| 29 | Onyx Pharmaceuticals | 6 | 1,578 | 9 | 8 |
| 30 | Whitehead Institute | 11 | 1,507 | 63 | 70 |
| 31 | Princeton University | 25 | 1,466 | 579 | 522 |
| 32 | Fred Hutchinson Cancer Research Center | 22 | 1,458 | 187 | 167 |
| 33 | University of Wisconsin–Madison | 19 | 1,406 | 2.1K | 1.2K |
| 34 | University of Georgia | 10 | 1,405 | 1.7K | 455 |
| 35 | Cornell University | 23 | 1,401 | 1.1K | 889 |
| 36 | Massachusetts Institute of Technology | 25 | 1,389 | 847 | 852 |
| 37 | University of Zurich | 15 | 1,375 | 1.0K | 504 |
| 38 | University of Rochester | 13 | 1,342 | 930 | 451 |
| 39 | Ludwig Cancer Research | 6 | 1,330 | 209 | 100 |
| 40 | Yale University | 19 | 1,302 | 977 | 744 |
| 41 | Duke University Hospital | 16 | 1,247 | 845 | 552 |
| 42 | Stanford University | 28 | 1,244 | 1.2K | 1.3K |
| 42 | Washington University School of Medicine | 20 | 1,244 | 713 | 548 |
| 44 | University of Toronto | 16 | 1,216 | 3.0K | 1.5K |
| 45 | University of Michigan | 14 | 1,215 | 2.9K | 1.6K |
| 46 | University of Pittsburgh | 12 | 1,200 | 2.1K | 919 |
| 47 | Michigan Medicine | 6 | 1,145 | 1.2K | 362 |
| 48 | Johns Hopkins University | 12 | 1,140 | 1.8K | 936 |
| 49 | St. Jude Children's Research Hospital | 7 | 1,130 | 376 | 149 |
| 50 | University of California, San Francisco | 18 | 1,119 | 741 | 557 |
| 51 | University of California, Berkeley | 13 | 1,105 | 775 | 538 |
| 52 | Brandeis University | 4 | 1,100 | 661 | 145 |
| 53 | University of North Carolina | 11 | 1,085 | 1.5K | 709 |
| 54 | Friedrich Miescher Institute for Biomedical Research | 9 | 1,080 | 51 | 46 |
| 55 | University of Iowa | 13 | 1,037 | 1.4K | 695 |
| 56 | Robert Wood Johnson Medical School | 5 | 1,031 | 372 | 71 |
| 57 | McGill University | 17 | 1,021 | 1.7K | 1.1K |
| 58 | Massachusetts General Hospital | 16 | 999 | 1.2K | 845 |
| 59 | Erasmus University Rotterdam | 3 | 975 | 1.7K | 287 |
| 60 | European Institute of Oncology | 5 | 965 | 238 | 77 |
| 61 | Northwestern University | 11 | 944 | 1.9K | 1.1K |
| 62 | Pennsylvania State University | 14 | 932 | 2.1K | 1.2K |
| 63 | University of British Columbia | 10 | 903 | 3.2K | 1.5K |
| 64 | University of Queensland | 7 | 888 | 3.4K | 1.2K |
| 65 | University of Arizona | 11 | 885 | 1.9K | 940 |
| 66 | Emory University | 13 | 852 | 1.1K | 702 |
| 67 | Eunice Kennedy Shriver National Institute of Child Health and Human Development | 11 | 836 | 271 | 220 |
| 68 | University of Texas at Austin | 9 | 823 | 3.4K | 1.6K |
| 69 | UC San Diego School of Medicine | 2 | 819 | 400 | 50 |
| 70 | University of Minnesota | 9 | 814 | 3.6K | 1.7K |
| 71 | Karolinska Institute | 6 | 811 | 2.2K | 956 |
| 72 | Dalhousie University | 4 | 800 | 2.3K | 453 |
| 73 | University of Tokyo | 9 | 796 | 3.5K | 1.4K |
| 74 | Canadian Institute for Advanced Research | 5 | 795 | 35 | 32 |
| 75 | Vita-Salute San Raffaele University | 2 | 791 | 471 | 64 |
| 76 | Hebrew University of Jerusalem | 8 | 790 | 1.5K | 561 |
| 77 | Trinity College Dublin | 5 | 789 | 1.0K | 255 |
| 78 | Lunenfeld-Tanenbaum Research Institute | 11 | 779 | 86 | 97 |
| 79 | Rutgers University | 8 | 777 | 2.2K | 937 |
| 80 | Stanford University Medical Center | 3 | 773 | 794 | 178 |
| 81 | Weizmann Institute of Science | 10 | 762 | 496 | 343 |
| 81 | University of Nottingham | 3 | 762 | 4.1K | 838 |
| 83 | Sapienza University of Rome | 4 | 739 | 3.3K | 605 |
| 83 | Kyoto University | 11 | 739 | 2.4K | 1.1K |
| 85 | Case Western Reserve University | 11 | 732 | 1.3K | 798 |
| 86 | Chinese Academy of Sciences | 11 | 715 | 3.5K | 1.9K |
| 87 | Michigan State University | 5 | 704 | 3.7K | 1.2K |
| 88 | Albert Einstein College of Medicine | 8 | 696 | 962 | 486 |
| 89 | Brown University | 10 | 671 | 1.1K | 717 |
| 90 | Imperial College London | 11 | 670 | 2.3K | 1.6K |
| 91 | Wellcome Sanger Institute | 13 | 658 | 85 | 112 |
| 92 | University of Basel | 5 | 646 | 1.5K | 555 |
| 92 | Wellcome Genome Campus | 9 | 646 | 95 | 113 |
| 94 | University of Maryland | 4 | 641 | 4.9K | 1.6K |
| 95 | Salk Institute for Biological Studies | 11 | 634 | 72 | 104 |
| 96 | Thomas Jefferson University | 5 | 627 | 1.3K | 442 |
| 97 | Heidelberg University | 12 | 617 | 1.3K | 982 |
| 97 | John Radcliffe Hospital | 14 | 617 | 291 | 334 |
| 99 | University of Virginia | 7 | 602 | 2.1K | 1.0K |
| 100 | Tel Aviv University | 3 | 585 | 4.4K | 1.1K |
| 101 | Free University of Brussels | 1 | 580 | 855 | 68 |
| 102 | University of Florida | 10 | 570 | 3.0K | 1.8K |
| 102 | Western General Hospital | 11 | 570 | 205 | 205 |
| 104 | Oregon Health & Science University | 1 | 568 | 2.8K | 518 |
| 105 | University of Southern California | 14 | 560 | 1.7K | 1.5K |
| 106 | Children's Hospital Boston | 4 | 553 | 319 | 188 |
| 107 | University of Oregon | 8 | 536 | 791 | 504 |
| 108 | National Human Genome Research Institute | 23 | 530 | 51 | 128 |
| 108 | École Normale Supérieure | 7 | 530 | 386 | 241 |
| 110 | University of Calgary | 2 | 514 | 5.5K | 1.2K |
| 111 | Nanjing Agricultural University | 1 | 503 | 1.6K | 281 |
| 112 | University of Cologne | 7 | 501 | 1.5K | 748 |
| 113 | Weill Cornell Medicine | 7 | 493 | 1.3K | 771 |
| 114 | ETH Zürich | 4 | 492 | 2.5K | 1.1K |
| 115 | Stowers Institute for Medical Research | 7 | 489 | 46 | 51 |
| 116 | University of Sheffield | 7 | 486 | 2.3K | 1.2K |
| 117 | University of Texas MD Anderson Cancer Center | 5 | 482 | 1.1K | 565 |
| 118 | Tufts University | 5 | 480 | 1.3K | 580 |
| 119 | Uppsala University | 5 | 477 | 2.9K | 1.3K |
| 120 | New York University | 12 | 457 | 1.1K | 1.1K |
| 121 | University of Copenhagen | 6 | 451 | 3.2K | 1.7K |
| 122 | Ohio State University | 3 | 448 | 6.4K | 2.2K |
| 123 | National Institute of Diabetes and Digestive and Kidney Diseases | 3 | 443 | 246 | 106 |
| 124 | Newcastle University | 6 | 442 | 1.9K | 950 |
| 125 | Vanderbilt University | 7 | 438 | 1.7K | 1.1K |
| 125 | Katholieke Universiteit Leuven | 2 | 438 | 5.4K | 1.5K |
| 127 | NYU Langone Medical Center | 1 | 432 | 1.7K | 338 |
| 128 | Indiana University Bloomington | 3 | 423 | 656 | 204 |
| 129 | Louisiana State University | 1 | 422 | 5.3K | 807 |
| 130 | Ajuntament de L’Hospitalet | 1 | 421 | 448 | 52 |
| 131 | National University of Singapore | 4 | 407 | 4.2K | 1.7K |
| 132 | German Cancer Research Center | 5 | 403 | 334 | 238 |
| 133 | University of Milan | 2 | 401 | 4.6K | 1.4K |
| 134 | Indiana University | 5 | 399 | 2.7K | 1.3K |
| 135 | Babraham Institute | 4 | 386 | 74 | 65 |
| 136 | University of Adelaide | 6 | 384 | 2.1K | 1.2K |
| 137 | Research Institute of Molecular Pathology | 4 | 382 | 44 | 47 |
| 138 | University of Manchester | 5 | 381 | 4.1K | 2.0K |
| 139 | National Institute of Genetics | 6 | 380 | 142 | 114 |
| 140 | Children's Medical Research Institute | 3 | 376 | 63 | 41 |
| 140 | Trinity College | 2 | 376 | 1.7K | 374 |
| 142 | QIMR Berghofer Medical Research Institute | 4 | 359 | 389 | 247 |
| 143 | Imperial College School of Medicine | 5 | 356 | 241 | 233 |
| 144 | University Medical Center Utrecht | 2 | 349 | 2.0K | 792 |
| 145 | Purdue University | 7 | 347 | 2.8K | 1.9K |
| 146 | University of Massachusetts | 4 | 345 | 2.5K | 1.2K |
| 147 | Duke University | 9 | 337 | 1.7K | 1.6K |
| 148 | University of Naples Federico II | 3 | 333 | 2.9K | 1.1K |
| 148 | Lawrence Berkeley National Laboratory | 5 | 333 | 742 | 592 |
| 150 | United States National Library of Medicine | 21 | 330 | 45 | 139 |
| 151 | University of Paris-Sud | 2 | 322 | 1.5K | 499 |
| 152 | Buck Institute for Research on Aging | 1 | 319 | 174 | 89 |
| 153 | University of North Carolina at Chapel Hill | 7 | 318 | 2.0K | 1.7K |
| 154 | University of Connecticut Health Center | 4 | 313 | 705 | 410 |
| 155 | University of Sydney | 3 | 311 | 6.0K | 2.6K |
| 155 | Geron Corporation | 1 | 311 | 37 | 30 |
| 157 | State University of New York Upstate Medical University | 2 | 310 | 1.1K | 279 |
| 158 | North Carolina State University | 5 | 307 | 2.5K | 1.5K |
| 159 | University of Lausanne | 5 | 298 | 1.2K | 816 |
| 160 | Université catholique de Louvain | 2 | 297 | 3.2K | 1.2K |
| 161 | J. Craig Venter Institute | 3 | 295 | 109 | 92 |
| 162 | Autonomous University of Madrid | 4 | 291 | 1.7K | 751 |
| 163 | Universidade Lusófona do Porto | 3 | 290 | 35 | 48 |
| 164 | University of Giessen | 2 | 289 | 2.3K | 794 |
| 165 | University of Utah | 6 | 280 | 2.7K | 1.9K |
| 166 | University of California, Los Angeles | 11 | 279 | 1.2K | 1.7K |
| 167 | University of Virginia Cancer Center | 1 | 273 | 50 | 9 |
| 168 | Commonwealth Scientific and Industrial Research Organisation | 2 | 271 | 1.4K | 609 |
| 169 | University of Melbourne | 5 | 270 | 4.6K | 2.8K |
| 169 | Colorado State University | 1 | 270 | 4.7K | 1.3K |
| 169 | Albany Medical Center | 1 | 270 | 1.2K | 272 |
| 172 | Natural History Museum | 1 | 269 | 1.1K | 320 |
| 173 | Monterey Bay Aquarium Research Institute | 1 | 263 | 160 | 60 |
| 174 | Stony Brook University | 5 | 260 | 1.7K | 1.2K |
| 175 | Huntsman Cancer Institute | 3 | 257 | 245 | 147 |
| 176 | European Molecular Biology Laboratory | 2 | 255 | 116 | 81 |
| 177 | Vanderbilt University Medical Center | 3 | 254 | 1.6K | 791 |
| 178 | University of Bath | 2 | 251 | 2.5K | 966 |
| 178 | Hammersmith Hospital | 3 | 251 | 838 | 516 |
| 180 | John Innes Centre | 7 | 249 | 165 | 218 |
| 181 | Technical University of Braunschweig | 1 | 248 | 2.0K | 423 |
| 182 | Columbia University Irving Medical Center | 8 | 246 | 748 | 648 |
| 183 | Osaka University | 8 | 244 | 2.0K | 1.5K |
| 183 | Cleveland Clinic | 4 | 244 | 1.9K | 1.2K |
| 185 | Collège de France | 5 | 242 | 200 | 201 |
| 186 | Paul Scherrer Institute | 1 | 241 | 983 | 330 |
| 187 | University of Leicester | 3 | 239 | 2.5K | 1.1K |
| 188 | University of Idaho | 1 | 236 | 2.1K | 503 |
| 189 | University of Colorado | 8 | 233 | 1.8K | 2.0K |
| 190 | National Heart, Lung, and Blood Institute | 4 | 230 | 534 | 433 |
| 191 | National Institute of Environmental Health Sciences | 3 | 228 | 622 | 429 |
| 192 | University of Birmingham | 2 | 227 | 5.5K | 2.2K |
| 193 | University of Vienna | 3 | 226 | 3.3K | 1.6K |
| 194 | Genome Institute of Singapore | 3 | 225 | 113 | 97 |
| 195 | University of Groningen | 1 | 218 | 5.9K | 2.1K |
| 196 | Iowa State University | 3 | 217 | 3.3K | 1.7K |
| 197 | University of Tsukuba | 2 | 214 | 3.6K | 1.2K |
| 197 | Carnegie Institution for Science | 5 | 214 | 246 | 283 |
| 199 | Erasmus MC | 3 | 212 | 1.7K | 1.0K |
| 199 | Anschutz Medical Campus | 4 | 212 | 948 | 755 |
| 201 | University of Kansas | 2 | 210 | 4.2K | 1.6K |
| 201 | University of Amsterdam | 2 | 210 | 5.3K | 2.5K |
| 203 | McGill University Health Centre | 1 | 209 | 1.9K | 670 |
| 203 | University of Warwick | 3 | 209 | 3.0K | 1.5K |
| 203 | Oslo University Hospital | 2 | 209 | 1.3K | 542 |
| 203 | University of California, Davis | 3 | 209 | 2.3K | 1.5K |
| 207 | Stockholm University | 2 | 207 | 3.1K | 1.4K |
| 208 | Dartmouth College | 2 | 206 | 1.8K | 822 |
| 209 | University of Glasgow | 4 | 205 | 3.2K | 2.0K |
| 210 | Florida State University | 3 | 204 | 2.7K | 1.4K |
| 211 | University of Newcastle | 2 | 201 | 3.6K | 1.5K |
| 212 | Goethe University Frankfurt | 4 | 199 | 1.6K | 1.1K |
| 212 | University of Alabama | 3 | 199 | 2.8K | 1.4K |
| 214 | PSL University | 5 | 198 | 409 | 299 |
| 215 | Mayo Clinic | 3 | 195 | 3.1K | 1.8K |
| 216 | Children's Hospital of Philadelphia | 6 | 194 | 809 | 731 |
| 217 | University of Fribourg | 1 | 192 | 1.6K | 430 |
| 217 | Austrian Academy of Sciences | 1 | 192 | 1.1K | 343 |
| 217 | University of South Carolina | 1 | 192 | 5.4K | 1.6K |
| 220 | University of Otago | 2 | 191 | 3.3K | 1.4K |
| 221 | Massey University | 2 | 186 | 2.0K | 796 |
| 222 | University of Sussex | 5 | 183 | 1.4K | 1.2K |
| 223 | University of Bern | 4 | 182 | 2.5K | 1.9K |
| 223 | Cardiff University | 4 | 182 | 2.4K | 1.7K |
| 223 | University of Manitoba | 1 | 182 | 5.6K | 1.7K |
| 226 | Australian National University | 5 | 181 | 2.4K | 1.9K |
| 227 | Leiden University Medical Center | 2 | 180 | 2.1K | 1.2K |
| 228 | China Medical University | 1 | 178 | 2.2K | 654 |
| 228 | University of Cincinnati Academic Health Center | 1 | 178 | 1.9K | 712 |
| 230 | École Normale Supérieure de Lyon | 7 | 176 | 244 | 337 |
| 231 | Curie Institute | 1 | 175 | 81 | 42 |
| 232 | Pompeu Fabra University | 3 | 174 | 715 | 427 |
| 233 | King's College London | 5 | 173 | 2.6K | 2.1K |
| 233 | University of Regensburg | 1 | 173 | 2.8K | 1.0K |
| 233 | The Christie NHS Foundation Trust | 2 | 173 | 288 | 178 |
| 236 | Ontario Cancer Institute | 3 | 171 | 190 | 207 |
| 237 | Washington State University | 3 | 166 | 2.8K | 1.8K |
| 237 | National Institute of Dental and Craniofacial Research | 1 | 166 | 625 | 358 |
| 239 | Nara Institute of Science and Technology | 2 | 162 | 600 | 256 |
| 240 | University of Helsinki | 6 | 161 | 2.6K | 2.6K |
| 241 | Radboud University Medical Centre | 2 | 159 | 1.2K | 821 |
| 242 | University of Bonn | 1 | 157 | 4.5K | 1.8K |
| 242 | University of California, Davis, MIND Institute | 1 | 157 | 116 | 55 |
| 244 | Tsinghua University | 3 | 156 | 3.5K | 2.1K |
| 244 | Arizona State University | 8 | 156 | 2.0K | 2.5K |
| 244 | Indiana University – Purdue University Indianapolis | 1 | 156 | 377 | 83 |
| 247 | NOVA University Lisbon | 2 | 154 | 1.7K | 744 |
| 248 | University of Tübingen | 2 | 153 | 3.4K | 1.8K |
| 249 | University of Chile | 1 | 152 | 4.2K | 1.2K |
| 249 | Université de Lyon | 5 | 152 | 750 | 754 |
| 251 | University of Oslo | 2 | 151 | 4.8K | 2.5K |
| 251 | Nagoya University | 3 | 151 | 2.9K | 1.7K |
| 251 | National Centre for Cell Science | 1 | 151 | 127 | 46 |
| 254 | French National Institute of Health and Medical Research | 4 | 144 | 1.7K | 1.4K |
| 255 | National Institute of Arthritis and Musculoskeletal and Skin Diseases | 1 | 143 | 250 | 163 |
| 256 | Agricultural Research Service | 1 | 142 | 1.2K | 688 |
| 257 | Tulane Medical Center | 1 | 137 | 287 | 98 |
| 258 | Federal University of Rio de Janeiro | 1 | 136 | 5.2K | 1.5K |
| 258 | Temple University | 1 | 136 | 4.4K | 1.5K |
| 260 | Russian Academy of Sciences | 3 | 135 | 3.9K | 2.1K |
| 261 | Leonard M. Miller School of Medicine | 3 | 134 | 1.7K | 1.3K |
| 262 | Jacobs University Bremen | 1 | 133 | 686 | 277 |
| 263 | University of Connecticut | 2 | 132 | 4.4K | 2.4K |
| 264 | University of Liège | 1 | 131 | 3.9K | 1.6K |
| 264 | Van Andel Institute | 1 | 131 | 195 | 127 |
| 266 | Children's Hospital Los Angeles | 1 | 130 | 969 | 449 |
| 267 | New York Blood Center | 1 | 128 | 239 | 133 |
| 268 | Houston Methodist Hospital | 3 | 127 | 585 | 468 |
| 269 | Medical University of Vienna | 2 | 125 | 2.2K | 1.4K |
| 270 | Addenbrooke's Hospital | 3 | 123 | 1.0K | 867 |
| 270 | Oakland University | 1 | 123 | 1.6K | 418 |
| 270 | Royal Melbourne Hospital | 1 | 123 | 1.1K | 527 |
| 273 | Federal University of Minas Gerais | 2 | 122 | 3.2K | 1.4K |
| 274 | University of Genoa | 1 | 121 | 4.3K | 1.7K |
| 275 | University of Texas Health Science Center at San Antonio | 1 | 120 | 2.2K | 1.1K |
| 276 | University of York | 3 | 119 | 2.3K | 1.7K |
| 277 | University of California, Irvine | 2 | 118 | 2.0K | 1.4K |
| 278 | Royal Netherlands Academy of Arts and Sciences | 2 | 116 | 76 | 104 |
| 279 | University of Antwerp | 1 | 114 | 3.9K | 1.6K |
| 280 | University of Ottawa | 3 | 113 | 3.8K | 2.8K |
| 281 | University of Illinois at Urbana–Champaign | 2 | 111 | 4.7K | 3.1K |
| 282 | Medical Research Council | 2 | 110 | 251 | 250 |
| 282 | The Graduate University for Advanced Studies | 2 | 110 | 277 | 173 |
| 284 | Charité | 2 | 107 | 1.4K | 971 |
| 285 | Miguel Hernández University of Elche | 1 | 106 | 1.3K | 477 |
| 285 | Clark University | 1 | 106 | 960 | 418 |
| 287 | Lille North of France University | 1 | 105 | 450 | 227 |
| 287 | Institut Pasteur de Lille | 1 | 105 | 459 | 290 |
| 287 | National Center for Agricultural Utilization Research | 1 | 105 | 282 | 176 |
| 287 | Wisconsin Energy Institute | 1 | 105 | 33 | 31 |
| 291 | Tongji University | 3 | 104 | 2.1K | 1.4K |
| 292 | University of Bergen | 3 | 101 | 2.3K | 2.1K |
| 292 | Spanish National Research Council | 4 | 101 | 643 | 864 |
| 294 | University of Barcelona | 1 | 99 | 6.1K | 2.8K |
| 294 | University of Brescia | 1 | 99 | 2.5K | 1.1K |
| 296 | TU Dresden | 3 | 98 | 2.2K | 1.8K |
| 296 | Joslin Diabetes Center | 1 | 98 | 175 | 154 |
| 298 | Université de Montréal | 4 | 97 | 3.1K | 3.1K |
| 298 | National Institute of Allergy and Infectious Diseases | 3 | 97 | 685 | 718 |
| 300 | Women's and Children's Hospital | 1 | 96 | 326 | 202 |
| 300 | New York State Department of Health | 1 | 96 | 853 | 506 |
| 302 | Regeneron Pharmaceuticals | 1 | 95 | 168 | 128 |
| 303 | University of Cincinnati | 1 | 94 | 5.3K | 2.5K |
| 303 | Oregon State University | 1 | 94 | 4.4K | 2.3K |
| 305 | Mississippi State University | 1 | 93 | 2.8K | 1.1K |
| 306 | Medical University of Graz | 1 | 92 | 1.8K | 941 |
| 306 | Wellcome Centre for Human Genetics | 1 | 92 | 64 | 61 |
| 308 | University of Nice Sophia Antipolis | 1 | 91 | 1.2K | 557 |
| 308 | UCL Medical School | 2 | 91 | 889 | 806 |
| 308 | New York State Psychiatric Institute | 1 | 91 | 744 | 500 |
| 311 | Assistance Publique – Hôpitaux de Paris | 1 | 88 | 1.7K | 960 |
| 312 | University of Texas at Dallas | 1 | 87 | 2.5K | 1.3K |
| 312 | Centre for Genomic Regulation | 1 | 87 | 39 | 40 |
| 314 | University of Bristol | 1 | 86 | 6.4K | 3.4K |
| 315 | Merck KGaA | 1 | 85 | 365 | 245 |
| 315 | University of East Anglia | 2 | 85 | 2.4K | 1.8K |
| 317 | University of Texas System | 1 | 83 | 2.4K | 1.4K |
| 318 | University of Illinois at Chicago | 2 | 82 | 4.3K | 3.1K |
| 319 | University of Perpignan | 1 | 81 | 253 | 121 |
| 319 | University of Guelph | 1 | 81 | 3.9K | 2.1K |
| 321 | University of Missouri | 1 | 79 | 6.7K | 3.5K |
| 322 | École normale supérieure Paris-Saclay | 1 | 77 | 651 | 320 |
| 323 | University of Wisconsin School of Medicine and Public Health | 1 | 76 | 998 | 669 |
| 324 | Shanghai Institute of Materia Medica | 1 | 75 | 482 | 321 |
| 325 | University of Liverpool | 3 | 74 | 3.8K | 3.4K |
| 325 | University Hospitals of Cleveland | 1 | 74 | 1.3K | 760 |
| 327 | University of Münster | 4 | 73 | 1.9K | 2.3K |
| 327 | University of Colorado Denver | 1 | 73 | 2.7K | 1.6K |
| 327 | Royal Botanic Gardens | 1 | 73 | 326 | 218 |
| 330 | Beatson West of Scotland Cancer Centre | 2 | 72 | 156 | 166 |
| 331 | Paris Diderot University | 1 | 70 | 2.2K | 1.3K |
| 331 | La Trobe University | 2 | 70 | 2.8K | 2.2K |
| 331 | (former) Institut National de Recherche pour l'Agriculture, l'Alimentation et l'Environnement | 3 | 70 | 1.5K | 1.7K |
| 331 | Developmental Biology Institute of Marseille | 2 | 70 | 35 | 44 |
| 335 | Marine Biological Association of the United Kingdom | 1 | 69 | 127 | 129 |
| 336 | Bielefeld University | 1 | 68 | 2.4K | 1.3K |
| 336 | Czech Academy of Sciences | 1 | 68 | 3.4K | 1.9K |
| 336 | Victoria University of Wellington | 1 | 68 | 2.7K | 1.2K |
| 336 | Malaghan Institute of Medical Research | 1 | 68 | 98 | 82 |
| 340 | Linköping University | 1 | 67 | 4.0K | 2.4K |
| 340 | Pontifical Catholic University of Chile | 1 | 67 | 3.1K | 1.5K |
| 342 | University of Hong Kong | 3 | 65 | 4.2K | 4.0K |
| 342 | Peking University | 2 | 65 | 4.3K | 3.3K |
| 342 | University of Central Lancashire | 1 | 65 | 1.5K | 707 |
| 342 | Necker–Enfants Malades Hospital | 1 | 65 | 707 | 487 |
| 346 | Guy's Hospital | 3 | 64 | 598 | 776 |
| 346 | Ulm University | 1 | 64 | 2.0K | 1.2K |
| 346 | Umeå University | 2 | 64 | 2.9K | 2.5K |
| 346 | Texas Children's Hospital | 1 | 64 | 921 | 592 |
| 350 | Humboldt University of Berlin | 2 | 63 | 3.4K | 2.7K |
| 350 | Niels Bohr Institute | 1 | 63 | 302 | 226 |
| 350 | RIKEN | 2 | 63 | 1.0K | 914 |
| 350 | National University of Ireland | 2 | 63 | 1.5K | 1.5K |
| 350 | Fujita Health University | 1 | 63 | 937 | 486 |
| 350 | Japan Science and Technology Agency | 1 | 63 | 962 | 690 |
| 356 | Pierre and Marie Curie University | 1 | 62 | 2.7K | 1.9K |
| 356 | University of Siena | 1 | 62 | 3.0K | 1.8K |
| 356 | Ottawa Hospital Research Institute | 1 | 62 | 934 | 695 |
| 359 | University of Lübeck | 1 | 61 | 1.3K | 868 |
| 360 | University of Freiburg | 1 | 60 | 3.9K | 2.5K |
| 360 | Université libre de Bruxelles | 2 | 60 | 2.9K | 2.5K |
| 360 | Academic Medical Center | 1 | 60 | 1.6K | 1.2K |
| 360 | Boston Medical Center | 1 | 60 | 1.2K | 784 |
| 360 | Royal Free Hospital | 1 | 60 | 1.2K | 745 |
| 360 | New York Genome Center | 2 | 60 | 38 | 57 |
| 366 | Stazione Zoologica Anton Dohrn | 3 | 59 | 123 | 194 |
| 366 | Sorbonne Universités | 2 | 59 | 866 | 883 |
| 368 | University of Bayreuth | 1 | 58 | 1.9K | 1.2K |
| 368 | University of New South Wales | 1 | 58 | 8.2K | 5.2K |
| 368 | London Research Institute | 2 | 58 | 104 | 172 |
| 371 | Macquarie University | 1 | 57 | 4.3K | 2.5K |
| 371 | Stellenbosch University | 1 | 57 | 3.5K | 2.0K |
| 371 | Western Sydney University | 1 | 57 | 2.8K | 1.7K |
| 371 | University of Alabama at Birmingham | 2 | 57 | 3.6K | 3.1K |
| 375 | Technical University of Munich | 3 | 56 | 3.6K | 3.8K |
| 376 | Carnegie Mellon University | 3 | 55 | 2.2K | 2.6K |
| 376 | University of Pavia | 3 | 55 | 2.3K | 2.5K |
| 378 | University of California, Santa Barbara | 2 | 54 | 1.5K | 1.6K |
| 379 | Technion – Israel Institute of Technology | 3 | 53 | 2.2K | 2.5K |
| 380 | Santa Fe Institute | 1 | 52 | 153 | 175 |
| 381 | Kookmin University | 1 | 50 | 741 | 413 |
| 381 | University of Bari | 1 | 50 | 2.9K | 2.0K |
| 383 | University of Oklahoma | 1 | 49 | 4.4K | 2.7K |
| 383 | Royal Holloway University of London | 1 | 49 | 918 | 655 |
| 385 | King's College | 1 | 48 | 2.5K | 1.7K |
| 386 | California Institute for Regenerative Medicine | 1 | 47 | 11 | 12 |
| 387 | Monash University | 2 | 46 | 6.2K | 5.4K |
| 388 | University of Strasbourg | 2 | 45 | 1.6K | 1.6K |
| 388 | University of Miami | 1 | 45 | 5.4K | 3.7K |
| 388 | Janelia Research Campus | 2 | 45 | 79 | 138 |
| 388 | University of Reading | 1 | 45 | 3.7K | 2.6K |
| 392 | Hungarian Academy of Sciences | 1 | 44 | 3.0K | 2.2K |
| 392 | Wageningen University & Research | 1 | 44 | 3.1K | 2.5K |
| 394 | McMaster University | 2 | 43 | 5.1K | 4.7K |
| 394 | Montana State University | 2 | 43 | 1.4K | 1.6K |
| 394 | Mellat Investment Bank | 1 | 43 | 7 | 9 |
| 397 | University of Hawaii | 1 | 41 | 3.0K | 2.3K |
| 397 | Paul Ehrlich Institute | 1 | 41 | 244 | 231 |
| 397 | University Hospital Würzburg | 1 | 41 | 559 | 447 |
| 397 | Oklahoma Medical Research Foundation | 1 | 41 | 395 | 389 |
| 397 | Sangamo Therapeutics | 2 | 41 | 18 | 47 |
| 402 | Tata Institute of Fundamental Research | 1 | 40 | 1.0K | 665 |
| 402 | National Centre for Biological Sciences | 1 | 40 | 245 | 220 |
| 402 | University of Montpellier | 2 | 40 | 1.6K | 1.7K |
| 402 | Helmholtz Zentrum München | 3 | 40 | 502 | 898 |
| 402 | University of Washington Friday Harbor Laboratories | 1 | 40 | 33 | 41 |
| 402 | Weatherall Institute of Molecular Medicine | 1 | 40 | 249 | 311 |
| 402 | Academia Sinica | 4 | 40 | 1.8K | 2.4K |
| 402 | Northeastern University | 1 | 40 | 5.0K | 3.5K |
| 410 | ComUE Paris-Saclay University | 2 | 39 | 2.1K | 1.8K |
| 410 | Simons Foundation | 1 | 39 | 22 | 37 |
| 412 | Leipzig University | 3 | 37 | 2.2K | 2.9K |
| 412 | Center for Research and Advanced Studies of the National Polytechnic Institute | 1 | 37 | 341 | 282 |
| 412 | Harwell Science and Innovation Campus | 1 | 37 | 362 | 304 |
| 415 | Great Ormond Street Hospital | 3 | 36 | 758 | 1.2K |
| 415 | Macdonald Campus | 1 | 36 | 137 | 142 |
| 415 | Montreal General Hospital | 1 | 36 | 612 | 606 |
| 415 | Université de Paris | 3 | 36 | 816 | 978 |
| 419 | Leiden University | 1 | 35 | 4.3K | 3.3K |
| 419 | Graduate University for Advanced Studies | 1 | 35 | 139 | 147 |
| 419 | Mount Saint Vincent University | 1 | 35 | 267 | 194 |
| 419 | UCSF Helen Diller Family Comprehensive Cancer Center | 1 | 35 | 75 | 107 |
| 423 | National Autonomous University of Mexico | 2 | 34 | 4.1K | 3.4K |
| 423 | Radboud University Nijmegen | 2 | 34 | 3.1K | 3.6K |
| 423 | Johns Hopkins Bloomberg School of Public Health | 1 | 34 | 2.1K | 1.9K |
| 423 | Georgia Institute of Technology | 3 | 34 | 2.7K | 3.7K |
| 423 | Pacific Northwest National Laboratory | 1 | 34 | 1.3K | 1.2K |
| 423 | Tufts Graduate School of Biomedical Sciences | 1 | 34 | 11 | 21 |
| 423 | National Institute for Basic Biology, Japan | 3 | 34 | 190 | 364 |
| 430 | Technical University of Madrid | 1 | 33 | 2.4K | 1.7K |
| 430 | Kansas State University | 1 | 33 | 3.5K | 2.7K |
| 430 | École polytechnique fédérale de Lausanne | 1 | 33 | 2.4K | 2.2K |
| 430 | University of New Brunswick | 1 | 33 | 2.1K | 1.5K |
| 434 | State University of New York | 2 | 32 | 4.9K | 5.1K |
| 434 | University at Albany | 2 | 32 | 1.0K | 1.4K |
| 436 | San Francisco General Hospital | 2 | 31 | 427 | 685 |
| 436 | Nicholas School of the Environment | 1 | 31 | 395 | 474 |
| 438 | Ocean University of China | 1 | 30 | 1.8K | 1.4K |
| 438 | Ehime University | 1 | 30 | 1.8K | 1.3K |
| 438 | St Mary's Hospital Medical School | 2 | 30 | 170 | 231 |
| 441 | Indian Statistical Institute | 1 | 29 | 1.1K | 785 |
| 441 | Claude Bernard University Lyon 1 | 2 | 29 | 1.9K | 2.3K |
| 443 | University of Alberta | 2 | 28 | 7.2K | 7.2K |
| 443 | University of Konstanz | 1 | 28 | 2.2K | 2.0K |
| 443 | Victor Chang Cardiac Research Institute | 2 | 28 | 101 | 211 |
| 443 | National Institute on Deafness and Other Communication Disorders | 1 | 28 | 185 | 236 |
| 443 | Institució Catalana de Recerca i Estudis Avançats | 1 | 28 | 371 | 502 |
| 448 | University of the Western Cape | 1 | 27 | 1.4K | 1.1K |
| 448 | Middle East Technical University | 1 | 27 | 2.5K | 2.0K |
| 448 | Brunel University | 1 | 27 | 2.2K | 2.1K |
| 448 | South African National Bioinformatics Institute | 1 | 27 | 45 | 61 |
| 448 | Sanford Consortium for Regenerative Medicine | 1 | 27 | 15 | 29 |
| 448 | Barcelona Biomedical Research Park | 1 | 27 | 32 | 76 |
| 448 | University of Colorado Anschutz Medical Campus | 2 | 27 | 799 | 1.1K |
| 455 | University of Warsaw | 1 | 26 | 3.3K | 2.2K |
| 455 | Lawrence Livermore National Laboratory | 1 | 26 | 1.5K | 1.5K |
| 457 | Nuffield Council on Bioethics | 1 | 25 | 13 | 14 |
| 457 | Université du Québec à Chicoutimi | 1 | 25 | 442 | 423 |
| 457 | University of Chinese Academy of Sciences | 3 | 25 | 2.0K | 2.6K |
| 457 | Okinawa Institute of Science and Technology | 1 | 25 | 367 | 351 |
| 461 | University of Exeter | 3 | 24 | 2.4K | 3.8K |
| 461 | Heriot-Watt University | 1 | 24 | 2.1K | 2.0K |
| 461 | James Hutton Institute | 1 | 24 | 413 | 486 |
| 461 | Sorbonne University | 2 | 24 | 1.6K | 1.7K |
| 465 | National Yang-Ming University | 1 | 23 | 2.0K | 1.9K |
| 465 | University of Tokushima | 1 | 23 | 1.7K | 1.3K |
| 465 | Instituto de Biomedicina y Biotecnología de Cantabria | 1 | 23 | 28 | 48 |
| 465 | Pacific Northwest Diabetes Research Institute | 1 | 23 | 80 | 130 |
| 465 | Agios Pharmaceuticals | 1 | 23 | 25 | 50 |
| 465 | Universidad de Cantabria | 1 | 23 | 878 | 802 |
| 471 | Oxford Brookes University | 2 | 22 | 983 | 1.5K |
| 471 | Utrecht University | 2 | 22 | 4.7K | 5.7K |
| 471 | Brigham Young University | 1 | 22 | 3.4K | 3.1K |
| 471 | University of Colorado Boulder | 1 | 22 | 3.2K | 3.1K |
| 471 | Swarthmore College | 1 | 22 | 553 | 553 |
| 471 | Centre hospitalier universitaire Sainte-Justine | 1 | 22 | 521 | 582 |
| 471 | Garvan Institute of Medical Research | 1 | 22 | 480 | 618 |
| 471 | Millersville University of Pennsylvania | 1 | 22 | 193 | 185 |
| 479 | University of the Witwatersrand | 1 | 21 | 4.2K | 3.5K |
| 479 | National Health Laboratory Service | 1 | 21 | 417 | 428 |
| 479 | Harvard Stem Cell Institute | 1 | 21 | 173 | 252 |
| 482 | Anhui University | 1 | 20 | 1.2K | 1.1K |
| 482 | Cambridge Biomedical Campus | 1 | 20 | 62 | 124 |
| 484 | University of Tartu | 2 | 19 | 1.7K | 2.4K |
| 484 | University of Aberdeen | 1 | 19 | 4.4K | 4.2K |
| 484 | Pablo de Olavide University | 2 | 19 | 461 | 779 |
| 484 | King's College Hospital | 1 | 19 | 1.3K | 1.3K |
| 484 | Estonian Biocentre | 2 | 19 | 16 | 57 |
| 489 | Lund University | 2 | 18 | 5.3K | 6.5K |
| 489 | Laval University | 1 | 18 | 5.2K | 4.9K |
| 489 | Universidad Europea de Madrid | 1 | 18 | 321 | 334 |
| 489 | Institut Jacques Monod | 1 | 18 | 158 | 248 |
| 489 | McGovern Institute for Brain Research | 1 | 18 | 159 | 244 |
| 489 | McKnight Brain Institute | 1 | 18 | 213 | 375 |
| 495 | Kumamoto University | 1 | 17 | 2.5K | 2.3K |
| 495 | Karolinska University Hospital | 1 | 17 | 2.4K | 2.5K |
| 495 | University of Gloucestershire | 1 | 17 | 357 | 429 |
| 498 | Tokyo Metropolitan University | 1 | 16 | 1.9K | 1.7K |
| 498 | Instituto Gulbenkian de Ciência | 1 | 16 | 281 | 402 |
| 498 | Transgene SA | 1 | 16 | 36 | 58 |
| 498 | Yale-NUS College | 1 | 16 | 122 | 167 |
| 502 | Autonomous University of Barcelona | 1 | 15 | 5.0K | 4.8K |
| 502 | Hiroshima University | 1 | 15 | 4.0K | 3.7K |
| 502 | NUI Galway | 1 | 15 | 1.4K | 1.7K |
| 502 | New York Medical College | 1 | 15 | 1.7K | 1.8K |
| 502 | Cancer Research Center | 1 | 15 | 265 | 415 |
| 502 | UNC Eshelman School of Pharmacy | 1 | 15 | 293 | 388 |
| 508 | Cold Spring Harbor Laboratory | 1 | 14 | 17 | 49 |
| 509 | Tohoku University | 3 | 13 | 3.4K | 4.9K |
| 509 | University of the Sciences | 1 | 13 | 255 | 395 |
| 511 | American Museum of Natural History | 1 | 12 | 550 | 668 |
| 511 | Waseda University | 1 | 12 | 3.0K | 2.9K |
| 511 | Miami University | 1 | 12 | 2.3K | 2.6K |
| 511 | University of São Paulo | 1 | 12 | 9.4K | 8.0K |
| 511 | Bethel University | 1 | 12 | 85 | 131 |
| 511 | Center for Infectious Disease Research | 1 | 12 | 110 | 216 |
| 511 | University of Minnesota Twin Cities | 1 | 12 | 471 | 860 |
| 518 | University of Kaiserslautern | 1 | 11 | 1.1K | 1.5K |
| 518 | Bar-Ilan University | 1 | 11 | 3.2K | 3.5K |
| 518 | PerkinElmer | 1 | 11 | 80 | 133 |
| 518 | University of Verona | 1 | 11 | 2.9K | 3.2K |
| 522 | Syracuse University | 1 | 10 | 3.4K | 3.9K |
| 522 | Tulane University | 1 | 10 | 3.2K | 3.8K |
| 522 | Stanley Center for Psychiatric Research at Broad Institute | 1 | 10 | 61 | 122 |
| 525 | European Bioinformatics Institute | 1 | 9 | 174 | 331 |
| 525 | University of Eastern Finland | 1 | 9 | 2.1K | 2.6K |
| 525 | University of Toronto Mississauga | 1 | 9 | 675 | 1.1K |
| 528 | University of Padua | 1 | 8 | 6.6K | 7.3K |
| 528 | Wayne State University | 1 | 8 | 5.2K | 5.9K |
| 528 | Hokkaido University | 1 | 8 | 5.1K | 5.3K |
| 528 | University Health Network | 1 | 8 | 2.1K | 2.7K |
| 528 | James Franck Institute | 1 | 8 | 104 | 177 |
| 528 | University of Nevada, Reno | 1 | 8 | 4.3K | 5.0K |
| 528 | University of Puerto Rico, Río Piedras Campus | 1 | 8 | 73 | 188 |
| 535 | University of London | 1 | 7 | 7.7K | 8.4K |
| 535 | Kyushu University | 1 | 7 | 5.6K | 6.0K |
| 535 | Aix-Marseille University | 1 | 7 | 3.0K | 3.8K |
| 538 | Tokyo Institute of Technology | 1 | 6 | 3.5K | 4.0K |
| 538 | AstraZeneca | 1 | 6 | 922 | 1.4K |
| 538 | University of Washington Medical Center | 1 | 6 | 660 | 1.1K |
| 541 | Amherst College | 1 | 5 | 478 | 774 |
| 541 | Konrad Lorenz Institute for Evolution and Cognition Research | 1 | 5 | 26 | 71 |
| 541 | University Hospital Heidelberg | 1 | 5 | 1.4K | 1.9K |
| 541 | University of Pretoria | 1 | 5 | 3.6K | 4.6K |
| 541 | University of Cape Town | 1 | 5 | 4.4K | 5.6K |
| 541 | University of Trieste | 1 | 5 | 2.8K | 3.9K |
| 541 | Human Technopole | 1 | 5 | 19 | 47 |
| 548 | National Tsing Hua University | 1 | 4 | 2.4K | 3.2K |
| 548 | University of Hohenheim | 1 | 4 | 1.5K | 2.3K |
| 548 | University of Florence | 1 | 4 | 5.1K | 6.5K |
| 548 | Southwest University | 1 | 4 | 2.2K | 2.9K |
| 548 | Aarhus University | 1 | 4 | 6.1K | 7.7K |
| 548 | University of Milan-Bicocca | 1 | 4 | 961 | 1.8K |
| 548 | Chongqing Normal University | 1 | 4 | 589 | 872 |
| 548 | Manchester Institute of Biotechnology | 1 | 4 | 121 | 281 |
| 556 | Michigan Technological University | 2 | 3 | 1.1K | 2.6K |
| 557 | Shanghai East Hospital | 1 | 2 | 482 | 920 |
| 557 | St George's, University of London | 1 | 2 | 916 | 1.5K |
| 559 | National Center for Biotechnology Information | 1 | 1 | 11 | 58 |
| 559 | Freie Universität Berlin | 1 | 1 | 4.4K | 6.4K |
| 561 | Amgen | 1 | 0 | 663 | 1.2K |
| 561 | University of Toulouse | 2 | 0 | 1.9K | 4.7K |
| 561 | University of Texas Health Science Center at Houston | 1 | 0 | 2.4K | 3.9K |
| 561 | Waksman Institute of Microbiology | 1 | 0 | 58 | 133 |
| 561 | MRC London Institute of Medical Sciences | 1 | 0 | 125 | 310 |
