| # | Institution | Articles | Citations | R ARank | R CRank |
|---|
| 1 | Howard Hughes Medical Institute | 70 | 39,587 | 100 | 31 |
| 2 | University of California | 78 | 32,060 | 2.0K | 238 |
| 3 | University of Edinburgh | 29 | 17,822 | 778 | 29 |
| 4 | University of Oxford | 47 | 15,994 | 780 | 99 |
| 5 | Yale University | 18 | 15,610 | 1.0K | 52 |
| 6 | Baylor College of Medicine | 34 | 15,410 | 526 | 48 |
| 7 | University of Washington | 31 | 14,546 | 1.4K | 142 |
| 8 | University of Cambridge | 45 | 13,538 | 801 | 122 |
| 9 | University of Chicago | 26 | 11,470 | 853 | 104 |
| 10 | Johns Hopkins University | 11 | 10,858 | 2.0K | 117 |
| 11 | Uppsala University | 18 | 10,143 | 1.0K | 50 |
| 12 | National Institutes of Health | 35 | 10,125 | 810 | 216 |
| 13 | Northwestern University | 7 | 9,708 | 2.7K | 115 |
| 14 | Friedrich Miescher Institute for Biomedical Research | 4 | 9,414 | 99 | 4 |
| 15 | Harvard University | 18 | 9,327 | 1.2K | 208 |
| 16 | Stanford University | 32 | 8,978 | 1.0K | 258 |
| 17 | University of Queensland | 17 | 8,888 | 1.7K | 61 |
| 18 | University of Southern California | 9 | 8,738 | 2.4K | 88 |
| 19 | Wellcome Genome Campus | 29 | 8,472 | 34 | 19 |
| 20 | University of California, San Diego | 19 | 8,230 | 871 | 112 |
| 21 | University of Texas at Austin | 17 | 8,225 | 2.1K | 158 |
| 22 | University of Notre Dame | 3 | 7,810 | 2.1K | 24 |
| 23 | National Center for Scientific Research | 20 | 7,746 | 2.4K | 260 |
| 24 | New York University | 11 | 7,733 | 1.2K | 74 |
| 25 | Wellcome Sanger Institute | 23 | 7,601 | 45 | 18 |
| 26 | Weizmann Institute of Science | 13 | 7,445 | 388 | 43 |
| 27 | Duke University Hospital | 11 | 7,272 | 1.1K | 115 |
| 28 | McGill University | 18 | 6,978 | 1.6K | 144 |
| 29 | The Rockefeller University | 12 | 6,887 | 307 | 56 |
| 30 | University College London | 18 | 6,875 | 1.6K | 188 |
| 31 | Fred Hutchinson Cancer Research Center | 14 | 6,816 | 273 | 58 |
| 32 | University of Michigan | 15 | 6,758 | 2.8K | 333 |
| 33 | Massachusetts General Hospital | 15 | 6,699 | 1.3K | 203 |
| 34 | University of California, Los Angeles | 8 | 6,362 | 1.6K | 102 |
| 35 | University of Oregon | 8 | 6,148 | 791 | 36 |
| 36 | University of British Columbia | 17 | 5,944 | 2.1K | 207 |
| 37 | Sidney Kimmel Comprehensive Cancer Center | 3 | 5,902 | 154 | 9 |
| 38 | Duke University | 12 | 5,796 | 1.3K | 124 |
| 39 | Pennsylvania State University | 14 | 5,308 | 2.1K | 188 |
| 40 | Memorial Sloan Kettering Cancer Center | 11 | 5,256 | 724 | 125 |
| 41 | University of Toronto | 17 | 5,241 | 2.9K | 404 |
| 42 | University of Copenhagen | 13 | 4,943 | 1.8K | 182 |
| 43 | USC Norris Comprehensive Cancer Center | 1 | 4,840 | 80 | 1 |
| 44 | Massachusetts Institute of Technology | 14 | 4,751 | 1.4K | 347 |
| 45 | University of Florida | 7 | 4,711 | 3.9K | 219 |
| 46 | University of Ottawa | 2 | 4,562 | 4.8K | 66 |
| 47 | Princeton University | 12 | 4,382 | 1.0K | 229 |
| 48 | University of Texas Southwestern Medical Center | 3 | 4,333 | 2.2K | 165 |
| 49 | Cornell University | 12 | 4,301 | 2.0K | 311 |
| 50 | Washington University School of Medicine | 9 | 4,229 | 1.3K | 217 |
| 51 | Newcastle University | 12 | 4,178 | 1.0K | 72 |
| 52 | University of Helsinki | 7 | 4,141 | 2.4K | 166 |
| 53 | Boston University | 6 | 4,120 | 1.9K | 127 |
| 54 | Karolinska Institute | 9 | 4,110 | 1.8K | 258 |
| 55 | Northeastern University | 1 | 3,987 | 5.0K | 75 |
| 56 | University of Georgia | 8 | 3,974 | 2.0K | 129 |
| 57 | Whitehead Institute | 12 | 3,949 | 58 | 32 |
| 58 | University of Sussex | 6 | 3,900 | 1.2K | 62 |
| 59 | University of Wisconsin–Madison | 10 | 3,858 | 3.3K | 459 |
| 60 | University of North Carolina | 6 | 3,820 | 2.5K | 226 |
| 61 | National Human Genome Research Institute | 13 | 3,814 | 91 | 32 |
| 62 | Imperial College London | 10 | 3,771 | 2.5K | 351 |
| 63 | North Carolina State University | 9 | 3,746 | 1.6K | 128 |
| 64 | University of Pennsylvania | 19 | 3,641 | 1.8K | 507 |
| 65 | Scripps Research | 7 | 3,621 | 521 | 131 |
| 66 | Indiana University | 10 | 3,604 | 1.5K | 156 |
| 67 | University of Manchester | 11 | 3,513 | 2.2K | 247 |
| 68 | Tel Aviv University | 9 | 3,363 | 2.2K | 160 |
| 69 | University of Rochester | 7 | 3,286 | 1.5K | 197 |
| 70 | Case Western Reserve University | 5 | 3,274 | 2.4K | 215 |
| 71 | Emory University | 6 | 3,198 | 2.0K | 213 |
| 72 | Michigan State University | 5 | 3,197 | 3.7K | 248 |
| 73 | Pasteur Institute | 7 | 3,169 | 595 | 107 |
| 74 | Washington State University | 5 | 3,064 | 2.0K | 107 |
| 75 | Ludwig Maximilian University of Munich | 4 | 3,062 | 3.0K | 232 |
| 76 | Australian National University | 3 | 3,039 | 3.3K | 177 |
| 76 | Georgia Institute of Technology | 4 | 3,039 | 2.3K | 207 |
| 78 | University of North Carolina at Chapel Hill | 4 | 2,996 | 3.0K | 251 |
| 79 | Garvan Institute of Medical Research | 3 | 2,983 | 233 | 28 |
| 80 | Columbia University | 9 | 2,959 | 3.0K | 522 |
| 80 | Aarhus University | 3 | 2,959 | 3.6K | 171 |
| 82 | King's Buildings | 2 | 2,896 | 179 | 5 |
| 83 | Beth Israel Deaconess Medical Center | 1 | 2,833 | 2.8K | 171 |
| 84 | Sangamo Therapeutics | 2 | 2,822 | 18 | 3 |
| 85 | NYU Langone Medical Center | 1 | 2,798 | 1.7K | 53 |
| 86 | University of Connecticut | 4 | 2,752 | 2.9K | 129 |
| 87 | QIMR Berghofer Medical Research Institute | 4 | 2,709 | 389 | 40 |
| 88 | Louisiana State University | 2 | 2,698 | 3.8K | 82 |
| 89 | University of California, Berkeley | 8 | 2,674 | 1.2K | 230 |
| 90 | University of Bath | 8 | 2,654 | 854 | 63 |
| 91 | Sapienza University of Rome | 2 | 2,631 | 4.8K | 127 |
| 92 | University of Melbourne | 10 | 2,551 | 2.8K | 445 |
| 93 | Babraham Institute | 2 | 2,526 | 133 | 7 |
| 94 | University of Montana | 4 | 2,520 | 520 | 36 |
| 95 | University of Arizona | 7 | 2,488 | 2.8K | 344 |
| 96 | King's College London | 10 | 2,421 | 1.5K | 273 |
| 97 | Chinese Academy of Sciences | 9 | 2,394 | 3.8K | 932 |
| 98 | University of Montpellier | 2 | 2,345 | 1.6K | 25 |
| 99 | Johns Hopkins Bloomberg School of Public Health | 5 | 2,343 | 891 | 120 |
| 100 | University of Maryland | 4 | 2,327 | 4.9K | 481 |
| 101 | Cardiff University | 4 | 2,326 | 2.4K | 166 |
| 102 | University of Utah | 6 | 2,312 | 2.7K | 303 |
| 103 | University of Zurich | 9 | 2,293 | 1.6K | 295 |
| 104 | Lawrence Berkeley National Laboratory | 4 | 2,277 | 840 | 177 |
| 105 | Joseph Fourier University | 3 | 2,263 | 522 | 40 |
| 106 | University of Sheffield | 5 | 2,251 | 2.9K | 286 |
| 107 | ETH Zürich | 6 | 2,239 | 2.0K | 324 |
| 108 | University of Idaho | 4 | 2,223 | 849 | 44 |
| 109 | Saarland University | 4 | 2,194 | 1.3K | 56 |
| 110 | University of Iowa | 3 | 2,190 | 4.0K | 346 |
| 111 | French National Institute of Health and Medical Research | 10 | 2,181 | 950 | 217 |
| 112 | University of Poitiers | 2 | 2,174 | 944 | 20 |
| 113 | University of Lausanne | 4 | 2,173 | 1.4K | 134 |
| 114 | St. Jude Children's Research Hospital | 3 | 2,170 | 669 | 94 |
| 115 | Brigham Young University | 2 | 2,085 | 2.3K | 66 |
| 116 | University of Würzburg | 3 | 2,062 | 2.1K | 191 |
| 117 | Victor Chang Cardiac Research Institute | 2 | 2,056 | 101 | 4 |
| 118 | University of Leicester | 6 | 2,042 | 1.4K | 167 |
| 119 | Wageningen University & Research | 7 | 2,023 | 1.1K | 283 |
| 120 | Trinity College Dublin | 4 | 1,998 | 1.3K | 82 |
| 121 | University of Groningen | 10 | 1,972 | 1.6K | 383 |
| 122 | University of Basel | 10 | 1,950 | 798 | 179 |
| 123 | St Bartholomew's Hospital | 1 | 1,923 | 628 | 26 |
| 124 | University of Vienna | 5 | 1,907 | 2.4K | 251 |
| 125 | Trinity College | 3 | 1,879 | 1.2K | 76 |
| 126 | University of London | 5 | 1,859 | 3.4K | 317 |
| 127 | Université de Montréal | 8 | 1,844 | 1.9K | 301 |
| 127 | University of Missouri | 3 | 1,844 | 4.0K | 240 |
| 129 | University of Konstanz | 3 | 1,822 | 1.2K | 85 |
| 130 | Helicos Biosciences | 1 | 1,770 | 11 | 2 |
| 131 | University Medical Center Utrecht | 6 | 1,756 | 1.1K | 243 |
| 132 | University of Sydney | 6 | 1,743 | 4.1K | 654 |
| 133 | Public Health England | 1 | 1,741 | 505 | 26 |
| 134 | Children's Hospital Boston | 1 | 1,728 | 781 | 60 |
| 135 | Albert Einstein College of Medicine | 6 | 1,719 | 1.2K | 208 |
| 136 | University of California, Davis | 7 | 1,714 | 1.2K | 224 |
| 137 | J. Craig Venter Institute | 5 | 1,688 | 69 | 32 |
| 138 | National Institute of Arthritis and Musculoskeletal and Skin Diseases | 1 | 1,671 | 250 | 28 |
| 139 | Temple University | 2 | 1,666 | 3.0K | 126 |
| 140 | Johns Hopkins School of Medicine | 4 | 1,665 | 1.4K | 232 |
| 141 | Western General Hospital | 7 | 1,657 | 296 | 80 |
| 142 | University of Minnesota | 6 | 1,644 | 4.8K | 982 |
| 142 | École polytechnique fédérale de Lausanne | 5 | 1,644 | 1.0K | 165 |
| 144 | University of Calgary | 3 | 1,632 | 4.5K | 417 |
| 145 | University of Pittsburgh | 3 | 1,630 | 5.2K | 702 |
| 146 | Vanderbilt University Medical Center | 4 | 1,623 | 1.4K | 172 |
| 147 | Sheba Medical Center | 2 | 1,616 | 977 | 49 |
| 148 | McMaster University | 2 | 1,599 | 5.1K | 457 |
| 149 | University of Adelaide | 3 | 1,580 | 3.4K | 288 |
| 150 | Vrije Universiteit Amsterdam | 4 | 1,556 | 2.3K | 319 |
| 151 | Technical University of Munich | 5 | 1,549 | 2.7K | 415 |
| 152 | Rutgers University | 6 | 1,542 | 2.8K | 462 |
| 153 | University of New Hampshire | 2 | 1,536 | 1.6K | 94 |
| 153 | Helmholtz Zentrum München | 6 | 1,536 | 283 | 52 |
| 155 | National Institute of Environmental Health Sciences | 6 | 1,515 | 382 | 114 |
| 156 | Austrian Academy of Sciences | 4 | 1,502 | 383 | 56 |
| 157 | Lunenfeld-Tanenbaum Research Institute | 2 | 1,501 | 347 | 55 |
| 158 | John Radcliffe Hospital | 4 | 1,481 | 822 | 163 |
| 159 | European Bioinformatics Institute | 8 | 1,462 | 44 | 26 |
| 160 | Children's Hospital of Philadelphia | 5 | 1,455 | 903 | 188 |
| 161 | Purdue University | 2 | 1,447 | 5.7K | 652 |
| 162 | Ghent University | 2 | 1,434 | 5.0K | 492 |
| 163 | Katholieke Universiteit Leuven | 11 | 1,345 | 1.7K | 590 |
| 164 | Mayo Clinic | 3 | 1,322 | 3.1K | 573 |
| 165 | Broad Institute | 7 | 1,309 | 44 | 35 |
| 165 | Academia Sinica | 1 | 1,309 | 3.5K | 203 |
| 167 | Gurdon Institute | 2 | 1,304 | 16 | 5 |
| 168 | Umeå University | 2 | 1,303 | 2.9K | 288 |
| 169 | Joint Genome Institute | 2 | 1,299 | 73 | 22 |
| 170 | University of Düsseldorf | 1 | 1,297 | 1.9K | 113 |
| 171 | University of Oulu | 2 | 1,294 | 2.5K | 147 |
| 172 | University of Pretoria | 1 | 1,288 | 3.6K | 90 |
| 173 | University of New South Wales | 2 | 1,271 | 6.4K | 706 |
| 174 | Addenbrooke's Hospital | 4 | 1,243 | 831 | 199 |
| 175 | Novo Nordisk Foundation Center for Protein Research | 2 | 1,227 | 17 | 2 |
| 176 | Florida State University | 2 | 1,220 | 3.4K | 326 |
| 177 | University of Houston | 1 | 1,205 | 4.5K | 219 |
| 178 | University of Tasmania | 3 | 1,194 | 1.8K | 133 |
| 179 | Ohio State University | 6 | 1,190 | 4.4K | 1.0K |
| 180 | University of Alabama | 1 | 1,186 | 5.1K | 244 |
| 181 | Graduate University for Advanced Studies | 1 | 1,179 | 139 | 6 |
| 181 | Centre for Genomic Regulation | 1 | 1,179 | 39 | 3 |
| 181 | Spanish National Research Council | 1 | 1,179 | 1.6K | 119 |
| 184 | University of California, Riverside | 1 | 1,177 | 1.7K | 88 |
| 185 | Heidelberg University | 4 | 1,171 | 2.9K | 598 |
| 186 | New York Genome Center | 2 | 1,160 | 38 | 9 |
| 187 | GlaxoSmithKline | 4 | 1,158 | 406 | 71 |
| 188 | Institute for Basic Science | 2 | 1,108 | 72 | 8 |
| 189 | Wayne State University | 3 | 1,107 | 3.0K | 328 |
| 190 | University of Tokyo | 4 | 1,105 | 5.3K | 1.1K |
| 190 | Swarthmore College | 4 | 1,105 | 162 | 18 |
| 192 | University of Texas MD Anderson Cancer Center | 3 | 1,092 | 1.5K | 309 |
| 193 | Norwegian University of Life Sciences | 1 | 1,070 | 1.2K | 75 |
| 194 | University of Exeter | 4 | 1,066 | 2.0K | 278 |
| 195 | California Institute of Technology | 5 | 1,061 | 1.4K | 570 |
| 196 | Massey University | 2 | 1,059 | 2.0K | 120 |
| 197 | University of Oslo | 2 | 1,052 | 4.8K | 573 |
| 198 | Erasmus University Rotterdam | 1 | 1,049 | 3.0K | 261 |
| 199 | Canadian Institute for Advanced Research | 1 | 1,038 | 124 | 26 |
| 200 | Oregon State University | 3 | 1,020 | 2.6K | 416 |
| 201 | University of Bristol | 6 | 1,019 | 2.6K | 737 |
| 202 | East Carolina University | 1 | 1,018 | 2.7K | 55 |
| 203 | University of KwaZulu-Natal | 2 | 1,017 | 1.8K | 70 |
| 204 | University of Birmingham | 6 | 1,006 | 2.9K | 711 |
| 204 | National Institute of Genetics | 2 | 1,006 | 296 | 67 |
| 204 | University of California, San Francisco | 8 | 1,006 | 1.4K | 604 |
| 204 | United States National Library of Medicine | 5 | 1,006 | 138 | 63 |
| 208 | Leiden University Medical Center | 5 | 995 | 1.4K | 451 |
| 209 | Hanyang University | 1 | 990 | 3.5K | 223 |
| 210 | The Roslin Institute | 3 | 987 | 105 | 15 |
| 211 | Erasmus MC | 2 | 983 | 2.0K | 389 |
| 212 | Ajuntament de L’Hospitalet | 2 | 966 | 251 | 20 |
| 213 | Salk Institute for Biological Studies | 3 | 962 | 199 | 75 |
| 213 | Columbia University Irving Medical Center | 5 | 962 | 1.0K | 240 |
| 215 | HUGO Gene Nomenclature Committee | 1 | 949 | 7 | 4 |
| 216 | German Cancer Research Center | 1 | 919 | 875 | 121 |
| 217 | Medical University of Graz | 1 | 912 | 1.8K | 138 |
| 218 | West Virginia University | 1 | 911 | 4.3K | 187 |
| 218 | Northern Arizona University | 1 | 911 | 1.6K | 83 |
| 218 | Evergreen State College | 1 | 911 | 105 | 3 |
| 221 | University of Illinois at Urbana–Champaign | 3 | 908 | 3.8K | 725 |
| 222 | Swedish University of Agricultural Sciences | 2 | 903 | 1.4K | 264 |
| 223 | National Institute on Alcohol Abuse and Alcoholism | 1 | 902 | 439 | 75 |
| 224 | University Hospitals of Cleveland | 2 | 899 | 873 | 118 |
| 225 | Van Andel Institute | 1 | 888 | 195 | 30 |
| 226 | Kaiser Permanente | 1 | 886 | 725 | 48 |
| 227 | University of California, Santa Cruz | 2 | 880 | 683 | 96 |
| 228 | University of Regensburg | 1 | 871 | 2.8K | 285 |
| 229 | University of Seville | 2 | 864 | 3.0K | 201 |
| 230 | Eunice Kennedy Shriver National Institute of Child Health and Human Development | 3 | 856 | 737 | 213 |
| 231 | National Institute on Aging | 3 | 854 | 510 | 188 |
| 232 | Eawag | 1 | 850 | 316 | 46 |
| 232 | University of Texas at Arlington | 1 | 850 | 2.4K | 110 |
| 234 | University of Oviedo | 1 | 846 | 2.8K | 134 |
| 235 | University of Colorado Boulder | 3 | 838 | 1.6K | 318 |
| 236 | University of Geneva | 4 | 811 | 1.8K | 438 |
| 237 | Ludwig Cancer Research | 4 | 809 | 290 | 138 |
| 238 | Radboud University Nijmegen | 3 | 805 | 2.5K | 506 |
| 239 | International Rice Research Institute | 1 | 804 | 370 | 62 |
| 239 | University of Reading | 4 | 804 | 1.7K | 365 |
| 241 | Weill Cornell Medicine | 3 | 786 | 2.1K | 548 |
| 242 | Kunming Institute of Zoology | 1 | 775 | 154 | 7 |
| 243 | Queensland Brain Institute | 2 | 770 | 151 | 24 |
| 244 | University of Alberta | 8 | 769 | 3.2K | 1.2K |
| 245 | Hebrew University of Jerusalem | 3 | 767 | 3.0K | 572 |
| 246 | University of Colorado | 4 | 753 | 3.0K | 837 |
| 246 | University of Maryland, Baltimore | 1 | 753 | 900 | 41 |
| 248 | The Graduate University for Advanced Studies | 1 | 752 | 434 | 23 |
| 249 | Carnegie Mellon University | 3 | 729 | 2.2K | 644 |
| 250 | China Agricultural University | 1 | 710 | 2.1K | 289 |
| 251 | Tufts Medical Center | 1 | 706 | 943 | 144 |
| 252 | Loyola University Chicago | 1 | 690 | 2.2K | 122 |
| 252 | University of Dayton | 2 | 690 | 672 | 49 |
| 254 | University of Nottingham | 5 | 687 | 3.0K | 905 |
| 255 | Carnegie Institution for Science | 4 | 682 | 281 | 147 |
| 256 | Charité | 2 | 676 | 1.4K | 232 |
| 257 | Utrecht University | 1 | 656 | 6.2K | 1.2K |
| 258 | Stowers Institute for Medical Research | 2 | 654 | 115 | 42 |
| 259 | University of St Andrews | 1 | 648 | 2.2K | 211 |
| 260 | Royal Netherlands Academy of Arts and Sciences | 1 | 645 | 114 | 34 |
| 261 | University of Tübingen | 3 | 636 | 2.8K | 695 |
| 262 | University of Veterinary Medicine Vienna | 1 | 635 | 469 | 42 |
| 263 | Pompeu Fabra University | 2 | 632 | 1.0K | 131 |
| 264 | Michigan Medicine | 2 | 624 | 2.3K | 576 |
| 265 | Mayo Clinic College of Medicine and Science | 1 | 606 | 1.5K | 286 |
| 266 | Montpellier 2 University | 2 | 605 | 390 | 64 |
| 267 | Jewish General Hospital | 2 | 604 | 527 | 81 |
| 268 | University of Cologne | 3 | 601 | 2.7K | 645 |
| 268 | Cooperative Institute for Research in Environmental Sciences | 1 | 601 | 288 | 82 |
| 270 | Wellcome Trust | 1 | 582 | 93 | 15 |
| 271 | University of Central Lancashire | 1 | 576 | 1.5K | 66 |
| 272 | Santa Fe Institute | 1 | 572 | 153 | 38 |
| 272 | Perinatal Institute | 1 | 572 | 76 | 11 |
| 274 | University of the Witwatersrand | 2 | 566 | 2.9K | 285 |
| 274 | University of Newcastle | 4 | 566 | 2.2K | 633 |
| 276 | Translational Genomics Research Institute | 1 | 558 | 199 | 37 |
| 277 | Swiss Institute of Bioinformatics | 1 | 556 | 170 | 40 |
| 278 | Stanford Cancer Institute | 1 | 554 | 96 | 25 |
| 279 | Warwick HRI | 1 | 553 | 176 | 42 |
| 280 | Monash University | 2 | 551 | 6.2K | 1.4K |
| 281 | Guy's Hospital | 3 | 550 | 598 | 199 |
| 282 | Hammersmith Hospital | 1 | 546 | 1.5K | 310 |
| 283 | University of Tennessee | 3 | 544 | 3.7K | 838 |
| 284 | Arizona State University | 3 | 542 | 4.0K | 1.1K |
| 285 | Agency for Science, Technology and Research | 2 | 540 | 388 | 99 |
| 286 | King's College | 1 | 534 | 2.5K | 281 |
| 287 | University of South Carolina | 1 | 533 | 5.4K | 728 |
| 287 | Tokyo Medical and Dental University | 1 | 533 | 2.3K | 353 |
| 289 | National University of Singapore | 1 | 530 | 7.7K | 1.4K |
| 289 | Hyogo College of Medicine | 1 | 530 | 1.2K | 99 |
| 291 | Dartmouth College | 3 | 529 | 1.3K | 384 |
| 292 | Leiden University | 3 | 525 | 2.4K | 713 |
| 293 | Cold Spring Harbor Laboratory | 2 | 520 | 10 | 9 |
| 294 | Iowa State University | 1 | 519 | 5.4K | 933 |
| 294 | Pacific Biosciences | 1 | 519 | 39 | 13 |
| 296 | Children's Medical Research Institute | 1 | 516 | 151 | 28 |
| 297 | Brown University | 3 | 515 | 2.8K | 865 |
| 298 | Tufts University | 1 | 514 | 3.6K | 540 |
| 299 | BC Centre for Disease Control | 1 | 505 | 149 | 14 |
| 300 | RWTH Aachen University | 5 | 502 | 1.9K | 650 |
| 301 | Johannes Gutenberg University Mainz | 1 | 501 | 2.8K | 442 |
| 302 | Medical College of Wisconsin | 2 | 500 | 2.1K | 513 |
| 303 | Osaka University | 1 | 496 | 5.3K | 956 |
| 304 | University of Hong Kong | 1 | 484 | 6.7K | 1.2K |
| 305 | Princess Alexandra Hospital | 1 | 459 | 1.0K | 141 |
| 306 | Food and Drug Administration | 2 | 458 | 601 | 129 |
| 306 | University of Delaware | 4 | 458 | 2.0K | 692 |
| 308 | National Tsing Hua University | 1 | 456 | 2.4K | 361 |
| 309 | Max Planck Institute for Demographic Research | 1 | 455 | 82 | 13 |
| 309 | University of Southern Denmark | 1 | 455 | 3.2K | 479 |
| 309 | National Institute of Diabetes and Digestive and Kidney Diseases | 1 | 455 | 501 | 103 |
| 312 | Cambridge University Hospitals NHS Foundation Trust | 2 | 452 | 500 | 105 |
| 313 | University of Warsaw | 1 | 451 | 3.3K | 267 |
| 313 | Inselspital | 1 | 451 | 696 | 83 |
| 315 | Francis Crick Institute | 1 | 443 | 88 | 16 |
| 316 | Macquarie University | 3 | 442 | 2.3K | 539 |
| 317 | University of Potsdam | 1 | 439 | 2.0K | 252 |
| 317 | University of Leeds | 5 | 439 | 3.1K | 1.3K |
| 319 | University of Bergen | 1 | 435 | 4.1K | 774 |
| 320 | Bio21 Institute | 1 | 434 | 128 | 17 |
| 321 | University of Barcelona | 1 | 433 | 6.1K | 1.1K |
| 322 | Lancaster University | 2 | 430 | 2.1K | 434 |
| 323 | University of Nevada, Reno | 1 | 427 | 4.3K | 465 |
| 324 | University of Côte d'Azur | 2 | 426 | 392 | 46 |
| 325 | École Normale Supérieure | 2 | 425 | 1.0K | 272 |
| 326 | Sanford Burnham Prebys Medical Discovery Institute | 1 | 424 | 420 | 126 |
| 327 | University of Amsterdam | 2 | 423 | 5.3K | 1.7K |
| 328 | Singapore Clinical Research Institute | 1 | 416 | 76 | 13 |
| 329 | Genome Institute of Singapore | 2 | 407 | 174 | 50 |
| 330 | Bar-Ilan University | 1 | 402 | 3.2K | 377 |
| 331 | European Molecular Biology Laboratory | 1 | 396 | 190 | 61 |
| 331 | Nagoya University | 1 | 396 | 4.5K | 881 |
| 333 | University of Bern | 2 | 385 | 3.7K | 1.2K |
| 333 | Huntsman Cancer Institute | 1 | 385 | 495 | 100 |
| 335 | University of Virginia | 2 | 380 | 4.7K | 1.4K |
| 336 | Harwell Science and Innovation Campus | 1 | 378 | 362 | 55 |
| 337 | Royal Devon and Exeter Hospital | 2 | 377 | 143 | 34 |
| 338 | University Hospital Heidelberg | 1 | 373 | 1.4K | 244 |
| 338 | Portland State University | 1 | 373 | 2.0K | 213 |
| 338 | Origins Institute | 1 | 373 | 16 | 3 |
| 341 | Medical University of Innsbruck | 2 | 364 | 1.1K | 333 |
| 342 | Humboldt University of Berlin | 1 | 362 | 4.7K | 885 |
| 343 | American Museum of Natural History | 1 | 354 | 550 | 122 |
| 343 | Colorado State University | 2 | 354 | 3.5K | 1.1K |
| 343 | Wildlife Conservation Society | 1 | 354 | 180 | 50 |
| 343 | Harvard Stem Cell Institute | 2 | 354 | 116 | 63 |
| 347 | Imperial College School of Medicine | 1 | 352 | 749 | 234 |
| 348 | University of California, Irvine | 2 | 351 | 2.0K | 650 |
| 349 | University of Freiburg | 2 | 350 | 2.8K | 890 |
| 350 | Ottawa Hospital Research Institute | 1 | 319 | 934 | 238 |
| 351 | University of Cape Town | 5 | 316 | 1.7K | 852 |
| 352 | Wellcome Trust Centre for Stem Cell Research | 1 | 309 | 11 | 7 |
| 353 | Biozentrum University of Basel | 1 | 305 | 107 | 44 |
| 354 | TU Dresden | 1 | 301 | 3.9K | 844 |
| 355 | Zhejiang A & F University | 1 | 300 | 214 | 15 |
| 356 | UCL Institute of Education | 1 | 298 | 401 | 38 |
| 357 | University of Southampton | 1 | 291 | 6.1K | 1.6K |
| 358 | University of Aberdeen | 1 | 286 | 4.4K | 1.1K |
| 358 | University of Stirling | 1 | 286 | 2.0K | 450 |
| 360 | University of Guelph | 1 | 281 | 3.9K | 985 |
| 360 | Anschutz Medical Campus | 2 | 281 | 1.4K | 621 |
| 362 | Wellcome Trust Centre for Cell-Matrix Research | 1 | 276 | 76 | 26 |
| 362 | Murdoch Children's Research Institute | 1 | 276 | 894 | 253 |
| 364 | Huazhong Agricultural University | 2 | 275 | 1.1K | 405 |
| 365 | University of Liverpool | 2 | 269 | 4.6K | 1.6K |
| 366 | Moscow State University | 1 | 268 | 3.8K | 743 |
| 367 | University of Liège | 1 | 262 | 3.9K | 1.0K |
| 368 | University of Alabama at Birmingham | 2 | 261 | 3.6K | 1.5K |
| 368 | VU University Medical Center | 3 | 261 | 1.1K | 691 |
| 370 | York University | 2 | 255 | 3.1K | 911 |
| 370 | National Institute on Deafness and Other Communication Disorders | 1 | 255 | 185 | 67 |
| 372 | Pablo de Olavide University | 1 | 253 | 825 | 75 |
| 373 | Oklahoma Medical Research Foundation | 1 | 251 | 395 | 142 |
| 374 | National University of General San Martín | 1 | 247 | 343 | 20 |
| 375 | University of Milan | 1 | 244 | 5.9K | 1.9K |
| 376 | Louis Pasteur University | 1 | 240 | 1.2K | 393 |
| 377 | University of South Australia | 1 | 236 | 3.1K | 675 |
| 377 | University of Nice Sophia Antipolis | 1 | 236 | 1.2K | 231 |
| 377 | Amsterdam University Medical Centers | 1 | 236 | 1.3K | 199 |
| 380 | Vanderbilt University | 3 | 234 | 3.1K | 1.6K |
| 381 | Stony Brook University | 1 | 233 | 4.3K | 1.3K |
| 381 | Royal Free Hospital | 1 | 233 | 1.2K | 329 |
| 383 | Boston College | 1 | 228 | 2.3K | 553 |
| 383 | Helmholtz Centre for Infection Research | 1 | 228 | 274 | 99 |
| 383 | Okinawa Institute of Science and Technology | 1 | 228 | 367 | 63 |
| 383 | Furness College | 1 | 228 | 16 | 1 |
| 387 | Simon Fraser University | 2 | 227 | 3.1K | 1.1K |
| 387 | Thomas Jefferson University | 3 | 227 | 1.8K | 951 |
| 389 | Rice University | 1 | 224 | 3.1K | 949 |
| 390 | University of Cincinnati | 1 | 223 | 5.3K | 1.4K |
| 390 | University of Lübeck | 1 | 223 | 1.3K | 364 |
| 390 | University of Zaragoza | 1 | 223 | 3.4K | 689 |
| 390 | Texas Tech University | 1 | 223 | 3.8K | 784 |
| 390 | University of Évry Val d'Essonne | 1 | 223 | 173 | 41 |
| 390 | John F. Kennedy Center for the Performing Arts | 1 | 223 | 32 | 6 |
| 390 | Medical University of Gdańsk | 1 | 223 | 1.3K | 219 |
| 390 | Greater Baltimore Medical Center | 1 | 223 | 47 | 15 |
| 390 | Kennedy Krieger Institute | 1 | 223 | 426 | 150 |
| 390 | Wills Eye Hospital | 1 | 223 | 189 | 48 |
| 390 | Royal Hospital for Sick Children, Edinburgh | 1 | 223 | 424 | 115 |
| 390 | Azienda Socio Sanitaria Territoriale Lariana | 1 | 223 | 37 | 2 |
| 402 | Oregon Health & Science University | 2 | 220 | 2.2K | 1.0K |
| 403 | Loyola University | 1 | 219 | 993 | 153 |
| 404 | Fudan University | 1 | 216 | 5.4K | 1.7K |
| 405 | Paris Diderot University | 1 | 215 | 2.2K | 630 |
| 405 | Sorbonne Paris Cité University (group) | 1 | 215 | 1.4K | 336 |
| 407 | Ontario Institute for Cancer Research | 2 | 214 | 98 | 69 |
| 407 | University of Western Australia | 2 | 214 | 4.0K | 1.8K |
| 409 | Kyoto University | 1 | 210 | 7.1K | 2.4K |
| 410 | John Innes Centre | 1 | 208 | 441 | 241 |
| 411 | Los Alamos National Laboratory | 1 | 206 | 2.0K | 815 |
| 412 | University of York | 2 | 204 | 2.9K | 1.2K |
| 413 | Lund University | 2 | 201 | 5.3K | 2.4K |
| 413 | Harvard-MIT Division of Health Sciences and Technology | 1 | 201 | 227 | 116 |
| 415 | Royal Institute of Technology | 1 | 196 | 3.1K | 972 |
| 415 | Churchill Hospital | 1 | 196 | 599 | 214 |
| 417 | Max Planck Institute for the Science of Human History | 1 | 195 | 84 | 23 |
| 418 | Beatson West of Scotland Cancer Centre | 1 | 187 | 228 | 90 |
| 419 | University of Hawaii | 1 | 186 | 3.0K | 941 |
| 420 | Prince of Wales Hospital | 1 | 176 | 1.8K | 625 |
| 420 | Northwell Health | 1 | 176 | 869 | 225 |
| 420 | The Feinstein Institute for Medical Research | 1 | 176 | 396 | 146 |
| 420 | Chinese University of Hong Kong | 1 | 176 | 5.9K | 2.1K |
| 424 | Institute for Research in Biomedicine | 1 | 169 | 56 | 38 |
| 424 | Aix-Marseille University | 2 | 169 | 2.1K | 839 |
| 426 | Weill Cornell Medical College in Qatar | 1 | 165 | 63 | 23 |
| 427 | University of Vigo | 1 | 163 | 1.7K | 435 |
| 427 | Icahn School of Medicine at Mount Sinai | 1 | 163 | 389 | 143 |
| 427 | National Heart, Lung, and Blood Institute | 1 | 163 | 1.2K | 524 |
| 430 | New York State Department of Health | 2 | 162 | 600 | 362 |
| 431 | University of Bonn | 1 | 156 | 4.5K | 1.8K |
| 432 | University of Dundee | 1 | 154 | 3.0K | 1.1K |
| 433 | University of Otago | 2 | 153 | 3.3K | 1.6K |
| 434 | Vita-Salute San Raffaele University | 1 | 151 | 747 | 281 |
| 434 | Genetic Alliance | 1 | 151 | 19 | 8 |
| 436 | University of Texas Health Science Center at Houston | 2 | 150 | 1.7K | 905 |
| 437 | McGill University Health Centre | 3 | 148 | 1.1K | 831 |
| 438 | University of Denver | 1 | 144 | 1.9K | 530 |
| 439 | Pohang University of Science and Technology | 1 | 143 | 1.4K | 565 |
| 440 | Andrés Bello University | 1 | 142 | 820 | 162 |
| 441 | University of Canterbury | 1 | 141 | 2.9K | 920 |
| 441 | University of Auckland | 1 | 141 | 6.3K | 2.4K |
| 441 | University of Waikato | 1 | 141 | 1.8K | 539 |
| 441 | South Australian Health and Medical Research Institute | 1 | 141 | 259 | 76 |
| 445 | University of Oklahoma | 3 | 139 | 2.3K | 1.5K |
| 445 | University Medical Center Groningen | 2 | 139 | 1.7K | 1.1K |
| 447 | IBM Thomas J. Watson Research Center | 1 | 137 | 514 | 267 |
| 447 | Sage Bionetworks | 1 | 137 | 59 | 42 |
| 449 | Czech Academy of Sciences | 1 | 134 | 3.4K | 1.4K |
| 450 | Radford University | 1 | 130 | 306 | 45 |
| 451 | University of Málaga | 2 | 126 | 1.8K | 773 |
| 452 | Maastricht University | 2 | 125 | 2.6K | 1.7K |
| 453 | Cranfield University | 1 | 124 | 1.6K | 720 |
| 453 | Carleton University | 1 | 124 | 3.5K | 1.2K |
| 453 | RIKEN | 1 | 124 | 1.5K | 623 |
| 453 | Memorial University of Newfoundland | 1 | 124 | 3.3K | 1.1K |
| 453 | Penn Genome Frontiers Institute | 1 | 124 | 8 | 12 |
| 458 | University of East Anglia | 2 | 120 | 2.4K | 1.5K |
| 459 | University of Toulouse | 1 | 119 | 2.8K | 1.1K |
| 460 | Texas Children's Hospital | 1 | 115 | 921 | 410 |
| 461 | Radboud University Medical Centre | 1 | 112 | 1.7K | 979 |
| 462 | Ben-Gurion University of the Negev | 2 | 109 | 3.3K | 1.8K |
| 462 | London School of Hygiene & Tropical Medicine | 1 | 109 | 274 | 159 |
| 464 | James Cook University | 1 | 108 | 2.3K | 939 |
| 465 | Oxford University Hospitals NHS Trust | 1 | 105 | 709 | 256 |
| 466 | Keck Graduate Institute | 2 | 103 | 45 | 59 |
| 467 | World Health Organization | 1 | 100 | 1.2K | 645 |
| 467 | National Autonomous University of Mexico | 2 | 100 | 4.1K | 1.9K |
| 467 | UCL Medical School | 1 | 100 | 1.2K | 779 |
| 470 | PAREXEL | 1 | 93 | 96 | 30 |
| 471 | University College Dublin | 1 | 92 | 4.2K | 1.9K |
| 472 | University of Turku | 1 | 91 | 4.1K | 2.0K |
| 473 | WEHI | 1 | 87 | 450 | 314 |
| 474 | Victoria University | 1 | 86 | 1.7K | 679 |
| 475 | PSL University | 1 | 85 | 1.3K | 552 |
| 476 | Natural History Museum | 1 | 84 | 1.1K | 637 |
| 477 | Delft University of Technology | 1 | 83 | 3.7K | 2.0K |
| 477 | Autism Speaks | 1 | 83 | 12 | 23 |
| 477 | University of Illinois at Chicago | 1 | 83 | 5.8K | 3.1K |
| 477 | Great Ormond Street Hospital | 1 | 83 | 1.4K | 855 |
| 477 | Kavli Institute of Nanoscience | 1 | 83 | 182 | 118 |
| 477 | Pennington Biomedical Research Center | 1 | 83 | 206 | 152 |
| 477 | Sorbonne University | 1 | 83 | 2.3K | 785 |
| 484 | University of Florence | 1 | 82 | 5.1K | 2.7K |
| 484 | University of Ferrara | 1 | 82 | 2.5K | 1.2K |
| 484 | Marche Polytechnic University | 1 | 82 | 2.4K | 1.2K |
| 487 | Dalhousie University | 1 | 81 | 5.0K | 2.6K |
| 487 | Laval University | 1 | 81 | 5.2K | 2.8K |
| 489 | National Institute of Allergy and Infectious Diseases | 1 | 80 | 1.1K | 788 |
| 490 | Wistar Institute | 1 | 79 | 426 | 326 |
| 491 | Queen Mary University of London | 1 | 78 | 2.5K | 1.3K |
| 492 | Rothamsted Research | 1 | 77 | 648 | 476 |
| 493 | University of Bayreuth | 1 | 76 | 1.9K | 1.0K |
| 494 | University College Cork | 1 | 75 | 2.7K | 1.4K |
| 495 | Bichat–Claude Bernard Hospital | 1 | 74 | 484 | 298 |
| 496 | Public Health Wales | 1 | 70 | 118 | 58 |
| 496 | Royal Veterinary College | 1 | 70 | 527 | 330 |
| 498 | University of Ljubljana | 1 | 69 | 4.1K | 2.0K |
| 498 | University of Luxembourg | 1 | 69 | 1.2K | 613 |
| 498 | University of Tartu | 1 | 69 | 2.5K | 1.3K |
| 498 | Technical University of Denmark | 1 | 69 | 3.4K | 2.3K |
| 498 | Flemish Institute for Technological Research | 1 | 69 | 197 | 146 |
| 498 | University of Ghana | 1 | 69 | 1.6K | 528 |
| 498 | University of Rwanda | 1 | 69 | 347 | 66 |
| 498 | Institute for Molecular Medicine Finland | 1 | 69 | 43 | 34 |
| 498 | Universidade do Porto | 1 | 69 | 2.4K | 1.2K |
| 507 | Technion – Israel Institute of Technology | 1 | 65 | 3.8K | 2.3K |
| 507 | University of Chinese Academy of Sciences | 1 | 65 | 3.2K | 1.8K |
| 509 | Toulouse III University - Paul-Sabatier | 1 | 60 | 485 | 352 |
| 510 | University of Trento | 2 | 57 | 1.6K | 1.4K |
| 511 | Beijing Institute of Genomics | 1 | 56 | 175 | 135 |
| 512 | Flinders University | 1 | 55 | 3.2K | 2.0K |
| 512 | City College of New York | 1 | 55 | 1.5K | 896 |
| 512 | University of Manitoba | 1 | 55 | 5.6K | 3.4K |
| 512 | Morton Arboretum | 1 | 55 | 11 | 13 |
| 512 | Centre d'Écologie Fonctionnelle et Évolutive | 1 | 55 | 164 | 112 |
| 512 | Federal Government of Belgium | 1 | 55 | 73 | 73 |
| 518 | National Centre for Biological Sciences | 2 | 54 | 161 | 179 |
| 518 | Cancer Research UK | 1 | 54 | 552 | 489 |
| 520 | Tata Institute of Fundamental Research | 1 | 53 | 1.0K | 572 |
| 520 | National University of Córdoba | 1 | 53 | 1.9K | 824 |
| 522 | George Mason University | 2 | 52 | 2.4K | 2.1K |
| 523 | Queen Elizabeth II Medical Centre | 1 | 46 | 143 | 145 |
| 523 | Harry Perkins Institute of Medical Research | 1 | 46 | 199 | 223 |
| 525 | Hungarian Academy of Sciences | 1 | 45 | 3.0K | 2.1K |
| 526 | University of Texas at Dallas | 1 | 40 | 2.5K | 1.9K |
| 526 | University of Chicago Medical Center | 2 | 40 | 460 | 542 |
| 528 | Public Health Agency of Canada | 1 | 39 | 394 | 391 |
| 528 | National Microbiology Laboratory | 1 | 39 | 123 | 131 |
| 530 | National University of Ireland | 1 | 37 | 2.4K | 2.0K |
| 530 | Bangalore University | 3 | 37 | 251 | 357 |
| 530 | Commonwealth Scientific and Industrial Research Organisation | 1 | 37 | 2.0K | 1.8K |
| 533 | Children's National Hospital | 1 | 35 | 1.1K | 967 |
| 534 | St. Michael's Hospital | 1 | 33 | 1.5K | 1.3K |
| 534 | University of Connecticut Health Center | 1 | 33 | 1.6K | 1.5K |
| 536 | University of Cincinnati Academic Health Center | 1 | 30 | 1.9K | 1.7K |
| 537 | University of Greifswald | 1 | 29 | 1.6K | 1.5K |
| 538 | National Institute of Standards and Technology | 1 | 26 | 1.9K | 1.7K |
| 538 | Ruhr University Bochum | 1 | 26 | 4.1K | 3.4K |
| 538 | Mitre Corporation | 1 | 26 | 321 | 263 |
| 541 | Royal Swedish Academy of Sciences | 1 | 24 | 91 | 140 |
| 541 | University of Hull | 1 | 24 | 2.7K | 2.4K |
| 543 | University of Patras | 2 | 20 | 2.0K | 2.7K |
| 543 | Claude Bernard University Lyon 1 | 1 | 20 | 2.6K | 2.6K |
| 543 | Université de Lyon | 1 | 20 | 2.3K | 2.3K |
| 546 | Cleveland Clinic | 1 | 18 | 3.3K | 3.1K |
| 546 | University Medical Center | 2 | 18 | 1.9K | 2.6K |
| 548 | State University of New York | 1 | 17 | 6.7K | 6.3K |
| 548 | BC Cancer Agency | 1 | 17 | 634 | 812 |
| 548 | Norwich Research Park | 1 | 17 | 220 | 353 |
| 551 | University of Antwerp | 2 | 14 | 2.7K | 4.1K |
| 552 | Colorado School of Public Health | 1 | 13 | 171 | 306 |
| 553 | Barcelona Biomedical Research Park | 1 | 12 | 32 | 100 |
| 554 | University of Vermont | 1 | 11 | 3.5K | 4.0K |
| 554 | Montpellier University Hospital | 1 | 11 | 341 | 477 |
| 556 | Virginia Tech | 1 | 10 | 5.9K | 6.2K |
| 557 | Southern Illinois University Carbondale | 1 | 9 | 3.2K | 3.6K |
| 558 | University of Massachusetts | 1 | 8 | 5.6K | 6.5K |
| 558 | University of Texas School of Law | 3 | 8 | 16 | 39 |
| 560 | University of Strathclyde | 1 | 7 | 3.8K | 4.7K |
| 560 | Royal College of Physicians and Surgeons of Canada | 1 | 7 | 71 | 143 |
| 560 | Elson S. Floyd College of Medicine | 1 | 7 | 90 | 184 |
| 563 | La Trobe University | 1 | 6 | 3.9K | 5.0K |
| 563 | University of Toronto Scarborough | 1 | 6 | 613 | 1.1K |
| 565 | University of Bologna | 1 | 5 | 6.9K | 8.0K |
| 565 | Birkbeck, University of London | 1 | 5 | 1.4K | 2.1K |
| 567 | Nationwide Children's Hospital | 1 | 4 | 826 | 1.3K |
| 568 | King Faisal Specialist Hospital and Research Centre | 1 | 3 | 476 | 829 |
| 568 | Muhimbili University of Health and Allied Sciences | 1 | 3 | 410 | 744 |
| 568 | Princess Anne Hospital | 1 | 3 | 114 | 232 |
| 571 | Stockholm University | 1 | 1 | 4.4K | 6.4K |
| 571 | Lehigh University | 1 | 1 | 2.1K | 3.4K |
| 571 | Stellenbosch University | 1 | 1 | 3.5K | 5.4K |
| 574 | International Livestock Research Institute | 1 | 0 | 271 | 568 |
| 574 | University of Otago, Christchurch | 1 | 0 | 198 | 449 |
